Six new species of the genus Metaphire Sims & Easton, 1972 (Annelida: Oligochaeta: Megascolecidae) from southeastern Vietnam
Author
Nguyen, Tung T.
Author
Nguyen, Nam Q.
Author
Lam, Dang H.
Author
Nguyen, Anh D.
text
Raffles Bulletin of Zoology
2020
2020-04-22
68
220
236
journal article
10.26107/RBZ-2020-0019
2345-7600
4577217
650E406A-6668-42DE-B6A6-637B154A10FE
Key to the species of
Metaphire
in southeastern
Vietnam
1. Male pores in xix ....................................................................2
– Male pores in xviii ..................................................................3
2. Spermathecal pores paired lateroventrally in 5/6/7/8/9. Clitellum normal, ring-shaped ..............
M. anomala
(
Michaelsen, 1907
)
– Spermathecal pores paired lateroventrally in 5/6/7/8. Clitellum saddle-shaped (
Fig. 6
) .........................
M. setosa
,
new species
3. Genital markings present in the male region .........................4
– Genital markings absent in the male region.........................12
4. Intestinal caeca manicate .......................................................... ....................................
M. pacseana
(
Thai & Samphon, 1988
)
– Intestinal caeca simple ............................................................5
5. Spermathecal pores in 5/6/7/8/9 .............................................6
– Spermathecal pores in 6/7/8/9 ................................................9
6. Genital markings in intersegment 19/20 and subsequent intersegments. Setal penis present ............................................ ..................................
M. malayanoides
Nguyen & Lam, 2017
– Genital markings present in segments. No setal penis ..........7
7. Only one pair of genital markings in xviii. Septum 8/9 absent ..................
M. grandiverticulata
Nguyen & Lam, 2017
– No genital markings in xviii. Septum 8/9 present ................8
8. Spermathecal pores located dorsally. Four to eight pairs of genital markings in xvii, xix, and subsequent segments (
Fig. 5
).................................................
M. songbeensis
,
new species
– Spermathecal pores located lateroventrally. Two pairs of genital markings in xvii and xix ..........
M. posthuma
(
Vaillant, 1868
)
9. Spermathecal pores located dorsally. Genital markings present in xvii and xix (
Fig. 2
)...........................
M. haui
,
new species
– Spermathecal pores located not dorsally. Genital markings present in intersegments 17/18 and 18/19............................10
10. Male region strongly concave..............
M. bahli
(
Gates, 1945
)
– Male region not concave. Genital markings disc-shaped ....11
11. The openings of copulatory pouches close to ventromedial line. Septum 10/11 absent ............
M. saigonensis
(
Omodeo, 1957
)
– The openings of copulatory pouches not close to ventromedial line. Septum 10/11 present .............
M. peguana
(
Rosa, 1890
)
12. Intestinal caeca manicate .....................................................13
– Intestinal caeca simple ..........................................................14
13. Spermathecal pores in 5/6/7/8/9. First dorsal pore in 11/12. Seminal vesicles connected (
Fig. 3
) ......................................... .......................................................
M. bariaensis
,
new species
– Spermathecal pores in 7/8/9. First dorsal pore in 10/11. Seminal vesicles separated ...................
M. californica
(
Kinberg, 1867
)
14. Spermathecal pore starting from 6/7 ....................................15
– Spermathecal pore starting from 7/8 ....................................17
15. Three spermathecal pores in 6/7/8/9......................................... ........................................................
M. houlleti
(
Perrier, 1872
)
– Two spermathecal pores in 6/7/8..........................................16
16. First dorsal pore in 12/13. Accessory glands present, ending at spermathecal ducts ...........
M. guillelmi
(
Michaelsen, 1895
)
– First dorsal pore in 11/12. Accessory glands absent (
Fig. 4
).. .....................................................
M. planatoides
,
new species
17. Two spermathecal pores in 7/8/9. No setal penis (
Fig. 7
) ...... ....................................................
M. houlletoides
,
new species
– Only one pair of spermathecal pores in 7/8. Setal penis present........................
M. xuanlocensis
Nguyen & Lam, 2017