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<mods:title id="A6A63EDBFA2D4496E7C56BB2661C6C6B">A review of the subgenus Pandinus Thorell, 1876 with descriptions of two new species from Uganda and Ethiopia (Scorpiones: Scorpionidae)</mods:title>
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(
<figureCitation id="13692A02A07D4778FE4CD11809248763" box="[439,554,1091,1114]" captionStart-0="Figures 1–5" captionStart-1="Figures 6–9" captionStart-10="Figure 42" captionStart-2="Figures 10–13" captionStart-3="Figures 14–16" captionStart-4="Figures 17–19" captionStart-5="Figures 20–22" captionStart-6="Figures 23–28" captionStart-7="Figure 29" captionStart-8="Figures 30–32" captionStart-9="Figures 33–41" captionStartId-0="3.[192,278,1871,1894]" captionStartId-1="5.[192,278,1860,1883]" captionStartId-10="16.[192,268,1821,1844]" captionStartId-2="6.[192,278,1846,1869]" captionStartId-3="7.[192,278,1786,1809]" captionStartId-4="9.[192,278,1832,1855]" captionStartId-5="10.[192,278,1862,1885]" captionStartId-6="12.[192,278,1846,1869]" captionStartId-7="13.[192,268,1230,1253]" captionStartId-8="14.[192,278,1856,1879]" captionStartId-9="15.[192,278,1768,1791]" captionTargetBox-0="[204,1394,193,1835]" captionTargetBox-1="[236,1405,167,1829]" captionTargetBox-10="[196,1437,169,1789]" captionTargetBox-2="[224,1401,197,1813]" captionTargetBox-3="[213,1417,167,1753]" captionTargetBox-4="[211,1417,176,1801]" captionTargetBox-5="[211,1417,167,1831]" captionTargetBox-6="[205,1424,193,1813]" captionTargetBox-7="[210,1424,195,1198]" captionTargetBox-8="[191,1438,159,1825]" captionTargetBox-9="[251,1403,194,1737]" captionTargetId-0="figure-49@3.[204,1408,192,1841]" captionTargetId-1="figure-6@5.[237,1404,168,1828]" captionTargetId-10="figure-14@16.[192,1440,165,1800]" captionTargetId-2="figure-10@6.[192,1437,192,1816]" captionTargetId-3="figure-107@7.[214,1416,168,1752]" captionTargetId-4="figure-6@9.[212,1416,177,1800]" captionTargetId-5="figure-14@10.[212,1416,168,1830]" captionTargetId-6="figure-14@12.[192,1440,192,1816]" captionTargetId-7="figure-6@13.[192,1440,192,1200]" captionTargetId-8="figure-14@14.[192,1437,160,1824]" captionTargetId-9="figure-61@15.[216,1418,191,1752]" captionTargetPageId-0="3" captionTargetPageId-1="5" captionTargetPageId-10="16" captionTargetPageId-2="6" captionTargetPageId-3="7" captionTargetPageId-4="9" captionTargetPageId-5="10" captionTargetPageId-6="12" captionTargetPageId-7="13" captionTargetPageId-8="14" captionTargetPageId-9="15" captionText-0="Figures 1–5: Pandinus mazuchi sp. n., ♀ (92.5 mm) holotype. 1–2. dorsal and ventral views. 3. Pectinal area. 4. Movable fin-" captionText-1="Figures 6–9: Pandinus mazuchi sp. n., ♀ holotype photographed at the type locality (6–7) and the type locality, Ethiopia, Jijiga env., 09°20'15.8&quot;N 42°42'17.5&quot;E, 2100 m a.s.l. (8–9)." captionText-10="Figure 42: Distribution map of the subgenus Pandinus. Localities of P. imperator and P. gambiensis are after Prendini (2004: 255, fig. 13), localities of other species are new records except Hargeysa (type locality of P. smithi). Type locality of P. phillipsii corresponds with new records (Goolis Mts.). Other localities of P. phillipsii and P. smithi cited by older authors (Borelli, 1919; Pocock, 1896, 1897, and 1900) are not included in the map. Exact distributions of these species need to be further investigated." captionText-2="Figures 10–13: Pandinus phillipsii (Pocock, 1896). 10–11. ♂ (125 mm), dorsal and ventral views, Somaliland, 15 km N of Sheikh, Goolis Mts., 09°58.927'N 45°10.377'E, 1247 m a.s.l. 12–13. ♀ (109 mm), dorsal and ventral views, Somaliland, Sheikh, 09°57'25.9&quot;N 45°09'52.2&quot;E, 1492 m a.s.l." captionText-3="Figures 14–16: Pandinus phillipsii (Pocock, 1896). Juvenile (80 mm) at the locality Somaliland, Sheikh, Goolis Mts., 09°56'23&quot;N 45°11'14.2&quot;E, 1439 m a.s.l. The large photo (14) is an overall view after overturning a smaller boulder. The smallest photo (15) shows a shallow depression under the boulder in which the scorpion was hiding and underwent an ecdysis (the same spot is shown in the large photo by an arrow). Small arrow points to a partial exuvia of this specimen. The lower photo (16) shows the entire juvenile after removal from the hiding place." captionText-4="Figures 17–19: Pandinus phillipsii (Pocock, 1896). 17–18. ♀ (109 mm) at the locality Somaliland, Sheikh, 09°57'25.9&quot;N 45°09'52.2&quot;E, 1492 m a.s.l. 19. ♂ (125 mm) on a tree after removal of dry bark, Somaliland, 15 km N of Sheikh, Goolis Mts., 09°58.927'N 45°10.377'E, 1247 m a.s.l." captionText-5="Figures 20–22: Pandinus smithi (Pocock, 1897), ♂ (107 mm), at the locality, Ethiopia, 55 km S of Degehbur, 07°49'27.2&quot;N 43°41'56.3&quot;E, 1053 m a.s.l. Red arrow points to the spot at which the specimen was found under bark." captionText-6="Figures 23–28: Pandinus ugandaensis sp. n. 23–25. ♂ (97 mm) holotype, dorsal and ventral views and pectinal area. 26–28. ♀ (105 mm) allotype, pectinal area and dorsal and ventral views." captionText-7="Figure 29: Pandinus ugandaensis sp. n., ♂ (97 mm) holotype, trichobothrial pattern. For explanation of trichobothrial nomenclature see Vachon (1974) and Kovařík (2009: 28, plate J)." captionText-8="Figures 30–32: Pandinus ugandaensis sp. n. 30. ♂ (97 mm) holotype. 31. ♀ (105 mm) allotype. 32. Uganda, Kaabong env., type locality and native collector." captionText-9="Figures 33–41: 33–36. Metasoma and telson in lateral views. 33. Pandinus ugandaensis sp. n., ♂ holotype. 34. Pandinus ugandaensis sp. n., ♀ allotype. 35. Pandinus mazuchi sp. n., ♀ holotype. 36. Pandinus imperator (C. L. Koch, 1841), ♂ (115 mm), Ghana, FKCP. 37–41. Tarsomere II of 4th leg. 37. Pandinus imperator (C. L. Koch, 1841), ♂ (115 mm), Ghana, FKCP. 38. Pandinus ugandaensis sp. n., ♂ holotype. 39. Pandinus mazuchi sp. n., ♀ holotype. 40. Pandinus phillipsii (Pocock, 1896), ♀ (109 mm), Somaliland, Sheikh, 09°57'25.9&quot;N 45°09'52.2&quot;E, 1492 m a.s.l., FKCP. 41. Pandinus smithi (Pocock, 1897), ♂ (107 mm), Ethiopia, 55 km of Degehbur, 07°49'27.2&quot;N 43°41'56.3&quot;E, 1053 m a.s.l., FKCP." figureDoi-0="http://doi.org/10.5281/zenodo.12782420" figureDoi-1="http://doi.org/10.5281/zenodo.12782424" figureDoi-10="http://doi.org/10.5281/zenodo.12782444" figureDoi-2="http://doi.org/10.5281/zenodo.12782426" figureDoi-3="http://doi.org/10.5281/zenodo.12782428" figureDoi-4="http://doi.org/10.5281/zenodo.12782430" figureDoi-5="http://doi.org/10.5281/zenodo.12782432" figureDoi-6="http://doi.org/10.5281/zenodo.12782436" figureDoi-7="http://doi.org/10.5281/zenodo.12782438" figureDoi-8="http://doi.org/10.5281/zenodo.12782440" figureDoi-9="http://doi.org/10.5281/zenodo.12782442" httpUri-0="https://zenodo.org/record/12782420/files/figure.png" httpUri-1="https://zenodo.org/record/12782424/files/figure.png" httpUri-10="https://zenodo.org/record/12782444/files/figure.png" httpUri-2="https://zenodo.org/record/12782426/files/figure.png" httpUri-3="https://zenodo.org/record/12782428/files/figure.png" httpUri-4="https://zenodo.org/record/12782430/files/figure.png" httpUri-5="https://zenodo.org/record/12782432/files/figure.png" httpUri-6="https://zenodo.org/record/12782436/files/figure.png" httpUri-7="https://zenodo.org/record/12782438/files/figure.png" httpUri-8="https://zenodo.org/record/12782440/files/figure.png" httpUri-9="https://zenodo.org/record/12782442/files/figure.png" pageId="2" pageNumber="1">Figs. 1–42</figureCitation>
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; 
<treatmentCitation id="0AF31096A07D4778FDDAD1DC081387A7" author="KRAEPELIN" box="[545,797,1159,1182]" page="116" pageId="2" pageNumber="1" year="1899">
<bibRefCitation id="EFC34B76A07D4778FDDAD1DC081387A7" author="KRAEPELIN" box="[545,797,1159,1182]" pageId="2" pageNumber="1" refId="ref7357" refString="KRAEPELIN, K. 1899. Scorpiones und Pedipalpi. In F. Dahl (ed.), Das Tierreich. Herausgegeben von der Deutschen Zoologischen Gesellschaft. Berlin: R. Friedlander und Sohn Verlag, 8. Lieferung. 265 pp." type="book" year="1899">Kraepelin, 1899: 116</bibRefCitation>
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; 
<treatmentCitation id="0AF31096A07D4778FF3BD1FE0A968785" author="VACHON" box="[192,408,1189,1212]" page="953" pageId="2" pageNumber="1" year="1974">
<bibRefCitation id="EFC34B76A07D4778FF3BD1FE0A968785" author="VACHON" box="[192,408,1189,1212]" pageId="2" pageNumber="1" pagination="857 - 958" refId="ref7847" refString="VACHON, M. 1974. Etude des caracteres utilises pour classer les familles et les genres de Scorpions. Bulletin du Museum national d´Histoire naturelle, Paris, 140: 857 - 958." type="journal article" year="1974">Vachon, 1974: 953</bibRefCitation>
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, figs. 113-118; 
<treatmentCitation id="0AF31096A07D4778FDB7D1FE08138785" author="SISSOM" box="[588,797,1189,1212]" page="136" pageId="2" pageNumber="1" year="1990">
<bibRefCitation id="EFC34B76A07D4778FDB7D1FE08138785" author="SISSOM" box="[588,797,1189,1212]" pageId="2" pageNumber="1" pagination="64 - 160" refId="ref7714" refString="SISSOM, W. D. 1990. Systematics, biogeography and paleontology. Pp. 64 - 160. In: Polis, G. A. (ed.): The Biology of Scorpions. Stanford: Stanford University Psress, 587 pp." type="book chapter" year="1990">Sissom, 1990: 136</bibRefCitation>
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; 
<treatmentCitation id="0AF31096A07D4778FF3BD19F0A6087E2" author="FET" box="[192,366,1220,1243]" page="248" pageId="2" pageNumber="1" year="1997">
<bibRefCitation id="EFC34B76A07D4778FF3BD19F0A6087E2" author="FET" box="[192,366,1220,1243]" pageId="2" pageNumber="1" pagination="245 - 250" refId="ref6958" refString="FET, V. 1997. Notes on the taxonomy of some Old World scorpions (Scorpiones: Buthidae, Chactidae, Ischnuridae, Scorpionidae). Journal of Arachnology, 25: 245 - 250." type="book chapter" year="1997">Fet, 1997: 248</bibRefCitation>
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; 
<treatmentCitation id="0AF31096A07D4778FE79D19F093E87E2" author="FET" box="[386,560,1220,1243]" page="465" pageId="2" pageNumber="1" year="2000">
<bibRefCitation id="EFC34B76A07D4778FE79D19F093E87E2" author="FET" box="[386,560,1220,1243]" pageId="2" pageNumber="1" refId="ref6995" refString="FET, V., W. D. SISSOM, G. LOWE &amp; M. E. BRAUNWALDER. 2000. Catalog of the Scorpions of the World (1758 - 1998). New York: The New York Entomological Society, 689 pp." type="book" year="2000">Fet, 2000: 465</bibRefCitation>
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; Prendini, 2000: 44; 
<treatmentCitation id="0AF31096A07D4778FF3BD1B80A8B87C3" author="KOVARIK" box="[192,389,1251,1274]" page="50" pageId="2" pageNumber="1" year="2009">
<bibRefCitation id="EFC34B76A07D4778FF3BD1B80A8B87C3" author="KOVARIK" box="[192,389,1251,1274]" pageId="2" pageNumber="1" refId="ref7185" refString="KOVARIK, F. 2009. Illustrated catalog of scorpions. Part I. Introductory remarks; keys to families and genera; subfamily Scorpioninae with keys to Heterometrus and Pandinus species. Prague: Clairon Production, 170 pp." type="book" year="2009">Kovařík, 2009: 50</bibRefCitation>
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, figs. 284-420.
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<taxonomicName id="4C524D04A07D4778FEA5D07D09D38604" authority="C. L. Koch, 1841" authorityName="C. L. Koch" authorityYear="1841" box="[350,733,1318,1341]" class="Arachnida" family="Buthidae" genus="Buthus" kingdom="Animalia" order="Scorpiones" pageId="2" pageNumber="1" phylum="Arthropoda" rank="species" species="imperator">
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C. L. 
<bibRefCitation id="EFC34B76A07D4778FDA6D07D09D38604" author="KOCH" box="[605,733,1318,1341]" pageId="2" pageNumber="1" pagination="695 - 698" refId="ref7043" refString="KOCH, C. L. 1841. Die Arachniden, 9 (1842): 108 pp. (Figs. 695 - 698), Nurnberg." type="book chapter" year="1841">Koch, 1841</bibRefCitation>
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.
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</subSubSection>
<subSubSection id="C348650CA07D4778FF3BD0310993854F" pageId="2" pageNumber="1" type="diagnosis">
<paragraph id="8BED3687A07D4778FF3BD0310993854F" blockId="2.[192,802,1386,1654]" pageId="2" pageNumber="1">
<smallCapsWord id="8D0BA05BA07D4778FF3BD0310A3886B9" baselines="1403,1403" box="[192,310,1386,1409]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Diagnosis" pageId="2" pageNumber="1">DIAGNOSIS</smallCapsWord>
. Total length 
<quantity id="4CAA9B62A07D4778FE25D031096686B8" box="[478,616,1386,1409]" metricMagnitude="-1" metricUnit="m" metricValue="1.4" metricValueMax="2.2" metricValueMin="0.6" pageId="2" pageNumber="1" unit="mm" value="140.0" valueMax="220.0" valueMin="60.0">60–220 mm</quantity>
. Pedipalp femur with 3 trichobothria, only one of them on internal surface. Pedipalp patella with 13–16 external and numerous (usually about 30) ventral trichobothria. Retrolateral pedal spurs absent. Lateroapical margins of tarsi produced into rounded lobes. Metasomal segments I–IV with paired ventral submedian carinae. Stridulatory organ located on opposing surfaces of pedipalp coxa and first leg. Telson without subaculear tubercle.
</paragraph>
</subSubSection>
</treatment>
</document>