A revision of the Palaearctic Pimeliini (Coleoptera: Tenebrionidae): a comparative analysis and systematic position of Eastern European and Asian taxa with dorso-lateral eyes Author Chigray, Svetlana N. 5097C803-AA1A-4856-95D7-F11DB747EB74 St. Petersburg State University, 16 line of Vasilevskiy Island, 29, St. Petersburg 199178 Russia. s.chigray@mail.ru Author Nabozhenko, Maxim V. A339C181-A65D-4F68-AA3E-01D07DFF25AD Precaspian Institute of Biological Resources of the Daghestan Federal Research Centre of the Russian Academy of Sciences, M. Gadzhiev str., 45, Makhachkala, Republic of Dagestan 367000 Russia. & Dagestan State University, M. Gadzhiev str., 43 a, Makhachkala, Republic of Dagestan 367000 Russia. & Severtsov Institute of ecology and evolution of the Russian Academy of sciences, Leninsky Prospect, 33, Moscow 119071 Russia. nalassus@mail.ru Author Chigray, Ivan A. 4CDA2EFB-5C20-4E11-870E-BDFD7DBB78F7 Zoological Institute of the Russian Academy of Sciences, Universitetskaya Emb. 1, St. Petersburg 199034 Russia. chigray93@bk.ru Author Abakumov, Evgeny V. 6E455A28-CEB1-419D-933D-98C9E9C2FED4 St. Petersburg State University, 16 line of Vasilevskiy Island, 29, St. Petersburg 199178 Russia. e.abakumov@spb.ru text European Journal of Taxonomy 2022 2022-04-05 809 1 71 http://dx.doi.org/10.5852/ejt.2022.809.1719 journal article 56065 10.5852/ejt.2022.809.1719 f76de735-01e3-454c-a7f1-bbafd26686f5 2118-9773 6419425 A754493E-5466-4479-B515-AABEDDE09D93 Genus Habrochiton Semenov Tjan-Shansky, 1907 Figs 1C–D , 10 , 15 , 26A , 27E Habrochiton Semenov-Tjan-Shansky, 1907: 179–181 . Type species Habrochiton vernus Semenov-Tjan-Shansky, 1907 , by original designation. Species included Habrochiton vernus Semenov-Tjan-Shansky, 1907 , H. primaeveris Semenov-Tjan-Shansky, 1907 . Diagnosis Body ( Fig. 1C–D ) brown, slender, elongate-oval, completely covered with moderately dense short ( H. vernus ) or long ( H. primaeveris ) longitudinally striated scales, sparse scale-like and simple setae ( Fig. 10 ). Body length 7–12 mm . Eyes small circular, moderately convex in dorsal view; surface behind eyes concealed under anterior margin of pronotum. Pronotum weakly transverse (1.4 × as wide as long). Anterolateral angles not expressed. Prosternum without transverse triangular depression along anterior margin. Prosternal process short and narrow Fig. 8. Apatopsis Semenov, 1891 and Habrobates Semenov, 1903 ,details of structure. A . A.grombczewskii Semenov, 1891 , ♂ (ZIN), pronotum. B . Ditto, scales and seta on pronotum. C . Ditto, elytral scales, setation and granulation. D . Ditto, granule, seta and scales on elytra. E . H. vernalis Semenov, 1903 , ♂, paralectotype (ZIN), pronotum. F . Scales and spinulose setation on pronotum. Fig. 9. Habrobates Semenov, 1903 and Dietomorpha Kühnelt, 1957 , details of structure. A . H. vernalis Semenov, 1903 , ♂, paralectotype (ZIN), left elytron. B . Ditto, elytral scales and rows of setae. C . D. pardalis Kühnelt, 1957 , ♂ (ZIN), pronotum. D . Ditto, scales, setae and granules in middle of pronotum. E . Ditto, base of left elytron. F . Ditto, scales, spine-like tubercle and seta on elytron. (length 1.7 × width), not raised above procoxae, not protruding. Prosternum very short, near 2 × as long as longitudinal diameter of one procoxa. Elytra elongate-oval ( H. vernus ) or triangular, almost wedge-shaped ( H. primaeveris ). Scutellar shield completely ( H. vernus ) or partially concealed ( H. primaeveris ) by elytral base; surface around scutellar shield without triangular depression. Transverse length of one metacoxa less than intercoxal process of abdominal ventrite 1 ( Fig. 1D ). Trochanters with very short dense setae. Meso- and metafemora curved outward along elytral lateral vertical side. Pro- and metatibiae weakly curved outward, protibiae ( Figs 26A , 27E ) widened at apex, outer margin of protibia projecting into short rounded process at apex with one ( H. primaeveris ) or several acutely angulate and closely located ( H. vernus ) short spines; outer margin armed with very short tubercles with long and thin spines and additionally with long and sparse setaesetae ( Fig. 27E ). Protibial terminal spurs strongly elongated, extending to base of protarsomere 5, mesotibial terminal spurs extending to base of mesotarsomere 3, metatibial terminal spurs extending to midlength of metatarsomere 1. Outer and inner protibial terminal spurs subequal in length, inner meso- and metatibial terminal spurs longer than outer one; spurs flattened, outer spur moderately narrowed to apex, acutely angulate at apex. Protarsi laterally not flattened, covered with short strong setae. Meso- and metatarsi flattened from sides, covered with long fine setae dorsally and shorter and stronger setae ventrally. Tarsal claws elongated, thin, weakly curved outward. Male genitalia Male genitalia identical to those in Apatopsis . Ovipositor ( Fig. 15J–L ) Very short, paraproct slightly longer than coxite. Paraproct with thin baculi, indistinct at apical third. Coxite lobe baculi short, more sclerotized in apical part, with additional sclerotized projection on ventral side; lobe II very narrow ( Fig. 15L ); lobe III slightly longer, conical, without additional projecting apical processes; lobe IV membranous, conical, rounded at apex, densely pubescent with long setae. Female genital ducts Destroyed during preparation. Distribution China (Taklamakan Desert).