A Revision of the Wasp Genus Pison Jurine, 1808 of Australia and New Zealand, New Guinea, and the Pacific Islands (Hymenoptera: Crabronidae) Author Pulawski, Wojciech J. text Proceedings of the California Academy of Sciences 2018 2018-12-17 65 1 584 journal article 10.5281/zenodo.13159946 0068-547X 13159946 Pison naralte Pulawski , species nova Figures 667 -674. NAME DERIVATION .– Naralte was an Australian aboriginal tribe that lived in the Renmark area, South Australia , where the holotype was collected; a noun in apposition to the generic name. RECOGNITION .– Pison naralte is an all black species, with three submarginal cells, the second recurrent vein interstitial with the second intersubmarginal vein or nearly so, and the setae appressed on tergum I. The female is unknown. The male is characterized by a non-emarginate sternum VIII combined with the presence of a tuft of erect setae on the apicolateral parts of sterna IV and V ( Fig. 670 ). Pison penicillatum is similar, although the erect setae are present on the apicolateral corner of sternum VII. Otherwise the two species differ as follows: in P. naralte the setae of the upper frons are about 0.5 × midocellar diameter long (about 1.0 × midocellar diameter in P. penicillatum ), those on the scutum are appressed (most P. penicillatum have sparse, erect setae whose length is at least one midocellar diameter), and those on the lower gena are shorter than the midocellar diameter (rather than 1.5-2.0 × as long as the midocellar diameter). Also, sternum II is closely punctate at least anterior of the apical depression (in P. penicillatum , sternum II is impunctate along the midline or has a few sparse punctures except closely punctate basally), sternum VIII has no longitudinal swelling (with obtuse, longitudinal swelling in P. penicillatum ), and erect setae of sternum V are 1.8-2.0 × as long as midocellar diameter (rather than as long as midocellar diameter on sternum VII). DESCRIPTION .– Frons dull, finely, shallowly punctate, punctures less than one diameter apart. Labrum not emarginate. Gena narrow in dorsal view ( Fig. 668 ). Anteromedian pronotal pit transversely elongate, about 2.5 × as long as midocellar diameter. Scutum not foveate along flange, without longitudinal ridges adjacent to posterior margin; scutal punctures fine but well defined, most punctures on disk averaging 2-3 diameter apart ( Fig. 669 ). Tegula enlarged. Mesopleural punctures compressed against each other or nearly so. Postspiracular carina present, about as long as midocellar diameter. Metapleural sulcus costulate between dorsal and ventral metapleural pits. Propodeum with irregular longitudinal carina separating side from dorsum and posterior surface and extending from gastral socket area toward spiracle; dorsum obliquely ridged in anterior half, transversely ridged in posterior half, punctate between ridges; side ridged, punctate between ridges; posterior surface transversely ridged, punctate between ridges. Hindcoxal dorsum with outer margin obtusely carinate. Outer surface of hindtibia with evanescent spines. Punctures of tergum I, anterior of apical depression, averaging slightly more than one diameter apart. Punctures of sternum II in holotype less than one diameter apart basally, up to two diameters apart on each side preapically and sublaterally, apical depression impunctate, in paratype closely punctate throughout. Setae silvery, nearly appressed on upper frons, appressed on postocellar area, scutum, and tergum I; on lower gena suberect, straight, shorter than midocellar diameter; largely concealing integument on clypeus (except lamella). Apical depressions of terga with silvery, setal fasciae. FIGURES 667-670. Pison naralte Pulawski , sp. nov. , male. (667) Clypeus and mandible; (668) Head in dorsal view; (669) Tegula and adjacent scutum; (670) Gastral apex in lateral view. FIGURES 671-673. Pison naralte Pulawski , sp. nov. , male. (671) Sternum VIII (ventral surface); (672) Genitalia in dorsal view; (673) Genitalia in lateral view. FIGURE 674. Collecting locality of Pison naralte Pulawski , sp. nov. Body all black, mandible brown mesally. ♀ .– Unknown ♂ .– Upper interocular distance equal to 0.80-0.84 × lower interocular distance; ocellocular distance equal to 1.2 × hindocellar diameter, distance between hindocelli equal to 0.9-1.3 × hindocellar diameter; eye height equal to 1.02 × distance between eye notches. Free margin of clypeal lamella acutely angulate ( Fig. 667 ). Dorsal length of flagellomere I 2.2 × apical width, of flagellomere X 1.3 × apical width. Sternum V with well-defined median sulcus on apical depression; apicolateral parts of sterna IV and V with tufts of erect setae ( Fig. 670 ), setae of sternum V 1.8-2.0 × as long as midocellar diameter. Apical margin of sternum VIII rounded ( Fig. 671 ). Genitalia: Figs. 672, 673 . Length 7.6-7.8 mm ; head width 2.3-2.4 mm . GEOGRAPHIC DISTRIBUTION (Fig. 674).– Known from a single locality in southeastern South Australia . RECORDS .– HOLOTYPE : ♂ , AUSTRALIA : South Australia : 79 km NNW Renmark at 33°31ˈS 140°24ˈE, 8 Nov – 12 Dec 1995 , K.R. Pullen ( ANIC ). PARATYPES : AUSTRALIA : South Australia : same data as holotype ( 1 ♂ , CAS ) ; same data as holotype except 12 Dec 1995 – 25 Jan 1996 ( 1 ♂ , ANIC ) .