The millipede genus Megaphyllum Verhoeff, 1894 in the Balkan Peninsula, with description of new species (Myriapoda: Diplopoda: Julida: Julidae)
Author
Lazányi, Eszter
Author
Vagalinski, Boyan
Author
Korsós, Zoltán
text
Zootaxa
2012
3228
1
47
journal article
45270
10.5281/zenodo.214875
6bae682e-cbd9-4e6a-a4af-e6c1fda188fb
1175-5326
214875
Megaphyllum margaritatum
(
Fanzago, 1875
)
Julus margaritatus
Fanzago, 1875
: 150
–151, no figures.
Chromatoiulus margaritatus
:
Attems 1927
: 239
–240, Figs 328–329;
Ceuca 1992
: 426
.
Chromatoiulus aetnensis
:
Attems 1927
: 224
.
Brachyiulus (Chromatoiulus) aetnensis
Verhoeff, 1910b
: 232
–236, Figs 31–32.
Brachyiulus (Italoiulus) garganensis
Verhoeff, 1932
: 490
–491,
Fig. 21
.
Brachyiulus (Italoiulus) aetnensis
:
Verhoeff 1932
: 491
.
Chromatoiulus (Italoiulus) garganensis
:
Attems 1940
: 308
.
Chromatoiulus (Italoiulus) aetnenis
:
Attems 1940
: 308
.
Chromatoiulus (Italoiulus) margaritatus
:
Attems 1940
: 308
;
Strasser 1974
: 290
, 296–297; 1976: 613.
Chromatoiulus (Italoiulus) margaritatus
var.
keratea
Strasser, 1974
: 277
, Figs 75–76.
Chromatoiulus (Italoiulus) margaritatus
var.
keratea
:
Strasser 1976
: 620
.
Chromatoiulus margaritatus kerateus
:
Ceuca 1992
: 426
.
Chromatoiulus (Italoiulus) margaritatus
var.
epirotica
Strasser, 1976
: 620
, Figs 63–65.
Chromatoiulus margaritatus epiroticus
:
Ceuca 1992
: 426
.
Megaphyllum margaritatum
:
Enghoff & Kime 2009
.
FIGURE 9.
Megaphyllum margaritatum
(Fanzago, 1875)
from Greece, Ossa Mts., male, right gonopod promere, caudal view (scale bar: 0.2mm). Abbreviation:
fl
: flagellum.
Material examined.
HNHM
:
Greece
: 233, 2ƤƤ, 2009/79, Larisa county, Ossa Mts., beech forest,
975m
,
N39°48.087’
E22°45.666’
, 0 9.04.2009, leg. LD, JK & DM.
Distribution in the Balkans.
Greece
: Attika: Keratea, Epirus: Elati (
Strasser 1974
,
1976
).
World distribution.
Widespread in
Italy
(
Foddai et al 1995
).
Remarks.
Although far from Keratea, the gonopods of the newly collected males resemble
var.
keratea
Strasser, 1974
. The flagella of the gonopod promeres of both males have a remarkable furcated apex (
fl
on
Fig. 9
).
Synonymy of
M. aetnense
,
M. garganense
and
M. margaritatum
has been discussed for a long time, especially by
Strasser. Strasser & Minelli (1984)
left the question open, writing that „…synonymy of
margaritatum
Fanz.
,
aetnense
Verh.
and
garganense
Verh.
is possible, but not sure…”. As we did not have enough individuals, we cannot add new information to this topic. However, if we accept the synonymy of these forms, the species seems to be very variable. In addition, the form
creticum
, one of the previous variants of
M. margaritatum
was elevated to species level several years ago (
Ćurčiċ et al 2001
). A thorough revision may produce similar results with other forms as well. Until then,
M. margaritatum
and
M. cretica
are the only two species in the subgenus
Italoiulus
.