Cochlostoma Jan, 1830 revised: an overview of the subgenus Turritus Westerlund, 1883 and its species (Caenogastropoda, Cochlostomatidae)
Author
Zallot, Enrico
365E17AD-6938-4364-A526-F1BDA2663E10
Per Ligont 1, 33070 Budoia, Italy.
ezallot@gmail.com
Author
Kamchev, Panche
9FA2BE01-3355-4340-B57C-9E2841D5E4C2
Naturalis Biodiversity Center, Darwinweg 2, 2333 CR Leiden, the Netherlands.
panekamchev@gmail.com
Author
Schilthuizen, Menno
683D0AB7-CDD8-4FAA-94B6-436F3BFB8873
Naturalis Biodiversity Center, Darwinweg 2, 2333 CR Leiden, the Netherlands. & Taxon Expeditions, Rembrandtstraat 20, 2311 VW Leiden, the Netherlands.
menno.schilthuizen@naturalis.nl
Author
Fehér, Zoltán
E801EC76-8B1E-450B-993E-BBBE57C00EA9
WWF Hungary, Álmos vezér útja 69 / A, 1141 Budapest, Hungary.
feher.zoltan.nhmus@gmail.com
Author
Mattia, Willy De
C74A049D-9D7B-4A1F-878B-5BDFD104219F
Central Research Laboratories, Burgring 7, 1010, Vienna, Austria; Department of Evolutionary Biology, University of Vienna, Djerassiplatz 1, 1030, Vienna, Austria.
wdemattia@gmail.com
Author
Gittenberger, Edmund
D786C279-FC92-4D08-AF16-F79A9705E0AE
Naturalis Biodiversity Center, P. O. Box 9517, 2300 RA Leiden, the Netherlands.
egittenberger@yahoo.com
text
European Journal of Taxonomy
2024
2024-03-21
927
1
163
https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2475/11013
journal article
10.5852/ejt.2024.927.2475
2118-9773
10853688
B6E43365-FACA-49BA-8CCD-77E4BF8F0016
Cochlostoma
(
T.
)
gracile croaticum
(
Pfeiffer, 1870
)
Figs 21
(cyan dots), 24–25
Pomatias croaticus
Pfeiffer, 1870: 38-15
–16.
Types
Not seen.
Other specimens
CROATIA
• 1-
Plitvice
(topotypical);
44.8640° N
,
15.5820° E
; 2002;
Murányi
leg.;
HNHM100120
•
2-
Korenica Lake
;
44.8931° N
,
15.6083° E
; 2000;
De Mattia
leg.;
WdM2175
•
3-
Gornja Ploča
,
Mount
Zir;
44.4305° N
,
15.6166° E
; 2017;
Fehér
leg.;
HNHM100626
•
4-
Velebit Mts
,
Prezid Pass
;
44.2477° N
,
15.8097° E
; 2005;
Murányi
leg.;
HNHM99895
.
Type locality
Pfeiffer (1870: 15–16)
listed several localities: “Habitat in
Croatia
: Trovera (Zelebor), Slunj, Brinj, Plitvice, Perusic, Klek (Brusina)”. Here, we select as locus typicus restrictus Plitvice.
Description
SHELL
. Widely spaced riblets on last part of protoconch. Teleoconch spotless, with moderately strong and rounded ribs becoming weaker approaching aperture. Moderately strong lip with columellar lobe abruptly inwardly curved to cover umbilicus.
MEASUREMENTS
.
6 ♀♀
: whorls =7.3–8.1, H =7.0–
7.8 mm
, H/W=2.59–2.76, roundness=0.14–0.19, ribs incl.= 59°–61°, apert. incl.= 14°–18°, ribs/mm 1
st
wh.=7–13, ribs/mm 4
th
wh.=8–16.
FEMALE
GENITAL
ORGANS
. As in
C.
(
T.
)
stussineri
.
Remarks
We did not amplify DNA of this taxon and therefore we provisionally report it as a subspecies of
C.
(
T.
)
gracile
. Of note is the different shell morphology of the male compared with the females, with much closer ribs (in this respect it is different from the male of
C.
(
T.
)
gracile gracile
from the
type
locality, which has ribs in the male as widely spaced as in the female), the darker color of the upper whorls and the sturdier shape, a sexual differentiation common in
Cochlostoma
.
Additional note to clade A
Within clade A there are then two supported branches, one with samples from the Central-Southern Appennine, which could be referred to
C.
(
T.
)
cassiniacum
(Saint-Simon in
Paulucci, 1878
) and
C.
(
T.
)
adamii
(
Paulucci, 1879
)
, and another with samples from
Albania
with
C.
(
T.
)
mnelense
(Wagner, 1914)
and a new species in the following described as
C.
(
T.
)
kontschani
sp. nov.
Beside these, there are polytomies in the BA or unsupported branches in the ML approach which include taxa from the Balkans: two are described entities from
Croatia
(
C.
(
T.
)
reitteri
(
Boettger, 1880
)
, currently reported as a subspecies of
C.
(
T.
)
gracile
) and
Greece
(
C.
(
T.
)
pageti
Klemm, 1962
); furthermore, there is a new species here described (
C.
(
T.
)
pallgergelyi
sp. nov.
) and two samples, one related to
C.
(
T.
)
pageti
and the other to the new species for which further studies are needed to understand their taxonomical status and here in the Appendix as NFS127 and NFS145. Finally, another sample from the Tarnova Forest in
Slovenia
, here in the Appendix as NFS158, raises issues which do not allow to properly classify it.