Cochlostoma Jan, 1830 revised: an overview of the subgenus Turritus Westerlund, 1883 and its species (Caenogastropoda, Cochlostomatidae) Author Zallot, Enrico 365E17AD-6938-4364-A526-F1BDA2663E10 Per Ligont 1, 33070 Budoia, Italy. ezallot@gmail.com Author Kamchev, Panche 9FA2BE01-3355-4340-B57C-9E2841D5E4C2 Naturalis Biodiversity Center, Darwinweg 2, 2333 CR Leiden, the Netherlands. panekamchev@gmail.com Author Schilthuizen, Menno 683D0AB7-CDD8-4FAA-94B6-436F3BFB8873 Naturalis Biodiversity Center, Darwinweg 2, 2333 CR Leiden, the Netherlands. & Taxon Expeditions, Rembrandtstraat 20, 2311 VW Leiden, the Netherlands. menno.schilthuizen@naturalis.nl Author Fehér, Zoltán E801EC76-8B1E-450B-993E-BBBE57C00EA9 WWF Hungary, Álmos vezér útja 69 / A, 1141 Budapest, Hungary. feher.zoltan.nhmus@gmail.com Author Mattia, Willy De C74A049D-9D7B-4A1F-878B-5BDFD104219F Central Research Laboratories, Burgring 7, 1010, Vienna, Austria; Department of Evolutionary Biology, University of Vienna, Djerassiplatz 1, 1030, Vienna, Austria. wdemattia@gmail.com Author Gittenberger, Edmund D786C279-FC92-4D08-AF16-F79A9705E0AE Naturalis Biodiversity Center, P. O. Box 9517, 2300 RA Leiden, the Netherlands. egittenberger@yahoo.com text European Journal of Taxonomy 2024 2024-03-21 927 1 163 https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2475/11013 journal article 10.5852/ejt.2024.927.2475 2118-9773 10853688 B6E43365-FACA-49BA-8CCD-77E4BF8F0016 Cochlostoma ( T. ) macei ( Bourguignat, 1870 ) Figs 105 (white dots), 111 Pomatias macei Bourguignat 1870: 56–57 . Types Not seen. Other specimens FRANCE • 1- Gourdon ; 43.7305° N , 6.959° E ; 1976; RMNH leg.; EZ0399 • 2- Gorge du Trevans ; 43.9471° N , 6.2267° E ; 2014; Zallot leg.; EZ1178 . Type locality FRANCE • Drôme, Grasse, Saint Vallier. Description SHELL . Closely spaced riblets on last part of protoconh. Teleoconch spotless with moderately strong ribs. Relatively strong lip with quite abruptly inwardly curved columellar lobe covering umbilicus. MEASUREMENTS . 3 ♀♀ : whorls= 8.3–9.1, H = 9.4–10.4 mm , H/W=2.79–3.1, roundness =0.1–0.14, ribs incl.= 57–59°, apert. incl.= 17–19°, ribs/mm 1 st wh.= 5–13, ribs/mm 4 th wh.=7–15. FEMALE GENITAL ORGANS . 2 specimens analysed. Slightly posterior connection of pedunculus to bursa copulatrix. Short, roughly oval seminal receptacle with undefined basis. 2–4 loops clustered beside apex of seminal receptacle. Junction of uterus gland far from connection of pedunculus of bursa copulatrix with distal oviduct. Remarks Our sample closest to the type locality originates from Gourdon, ca 10 km east of Saint Vallier. The other sample, from the Gorge du Trevans, is located 50 km northeast of the type locality. Only of the latter we were able to analyse the genitals. The shell morphology and the female genitalia suggest that this species belongs to Turritus . Only H3 was sequenced and the results support this conclusion. Based on some similarity of the female genital morphology with that of C. ( T. ) sardoum , it could belong to the same subclade within clade B. This would be interesting in a paleogographic perspective because it would be the ‘remnant’ on the European plate of the closest relative to the sardoum , agriotes / affine , and atlanticum taxa (see Discussion), which originated on the southern border of the European plates before the geological eras which led to the current shape of the western Mediterranean Sea and the lands that sourround it.