Cochlostoma Jan, 1830 revised: an overview of the subgenus Turritus Westerlund, 1883 and its species (Caenogastropoda, Cochlostomatidae)
Author
Zallot, Enrico
365E17AD-6938-4364-A526-F1BDA2663E10
Per Ligont 1, 33070 Budoia, Italy.
ezallot@gmail.com
Author
Kamchev, Panche
9FA2BE01-3355-4340-B57C-9E2841D5E4C2
Naturalis Biodiversity Center, Darwinweg 2, 2333 CR Leiden, the Netherlands.
panekamchev@gmail.com
Author
Schilthuizen, Menno
683D0AB7-CDD8-4FAA-94B6-436F3BFB8873
Naturalis Biodiversity Center, Darwinweg 2, 2333 CR Leiden, the Netherlands. & Taxon Expeditions, Rembrandtstraat 20, 2311 VW Leiden, the Netherlands.
menno.schilthuizen@naturalis.nl
Author
Fehér, Zoltán
E801EC76-8B1E-450B-993E-BBBE57C00EA9
WWF Hungary, Álmos vezér útja 69 / A, 1141 Budapest, Hungary.
feher.zoltan.nhmus@gmail.com
Author
Mattia, Willy De
C74A049D-9D7B-4A1F-878B-5BDFD104219F
Central Research Laboratories, Burgring 7, 1010, Vienna, Austria; Department of Evolutionary Biology, University of Vienna, Djerassiplatz 1, 1030, Vienna, Austria.
wdemattia@gmail.com
Author
Gittenberger, Edmund
D786C279-FC92-4D08-AF16-F79A9705E0AE
Naturalis Biodiversity Center, P. O. Box 9517, 2300 RA Leiden, the Netherlands.
egittenberger@yahoo.com
text
European Journal of Taxonomy
2024
2024-03-21
927
1
163
https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2475/11013
journal article
10.5852/ejt.2024.927.2475
2118-9773
10853688
B6E43365-FACA-49BA-8CCD-77E4BF8F0016
Cochlostoma
(
T.
)
simrothi
(
Caziot, 1908
)
Figs 105
(magenta dot), 106
Pomatias simrothi
Caziot, 1908: 460
.
Types
Not seen.
Other specimen
FRANCE
• 1-
Saorge
(topotypical);
43.9881° N
,
7.5513° E
; 2009;
Margelli
leg.;
EZ0782
.
Type locality
FRANCE
•
Provence-Alpes-Côte d’Azur
, Gorge de Saorge.
Description
SHELL
. Last part of protoconch ribbed with very closely spaced riblets. Teleoconch spotless, with moderately strong and rounded ribs becoming weaker approaching aperture. Strong and developed lip. Columellar lobe bent inwardly to cover umbilicus.
Fig. 105.
Distribution of samples with undetermined position in the phylogenetic trees: white=
Cochlostoma
(
T.
)
macei
(
Bourguignat, 1870
)
; purple =
C.
(
T.
)
sturanii
(
Wagner, 1897
)
; orange=
C.
(
T.
)
kleciaki
(
Braun, 1887
)
; yellow=
C.
(
T.
)
arnautorum
(
Wagner, 1906
)
; red =
C.
(
T.
)
atlanticum
(Bourguignat, 1868)
; green =
C.
(
T.
)
euboicum
(
Westerlund, 1885
)
; blue =
C.
(
T.
)
martensianum
(
Möllendorff, 1873
)
; magenta=
C.
(
T.
)
simrothi
(
Caziot, 1908
)
; cyan =
C.
(
T.
)
fontqueri
Haas, 1924
.
MEASUREMENTS
.
2 ♀♀
: whorls =8.1–8.2, H =
8.1–8.8 mm
, H/W= 2.62–2.76, roundness =0.1–0.11, ribs incl.= 53–60°, apert. incl.= 14–15°, ribs/mm 1
st
wh.= 8–8, ribs/mm 4
th
wh.=8–10.
FEMALE
GENITAL
ORGANS
(
1 ♀
). Ventral connection of pedunculus of bursa copulatrix. One or no loops between apex of seminal receptacle and proximal loop.Almost tubular seminal receptacle with undefined basis. Junction of uterus gland close to connection between pedunculus of bursa copulatrix and distal oviduct.
Fig. 106.
Topotypical
Cochlostoma
(
T.
)
simrothi
(
Caziot, 1908
)
, 1- Saorge, FR (EZ-0782).
Remarks
Cochlostoma
(
T.
)
simrothi
lives in the Maritime Alps, an area where different taxa have been described (
Pomatias simrothi
Caziot, 1908
,
Pomatias acutum
Caziot, 1908
,
Pomatias galloprovincialis
Locard, 1894
and
Pomatias subalpinus
Pini, 1885
). We lack a proper sampling of this area which needs a dedicated study.
Previous authors considered
C.
(
T.
)
simrothi
a subspecies of
C.
(
T.
)
patulum
(
Zilch 1958
; “possibly conspecific” in
Welter-Schultes 2012: 101
). The similarities of the shell suggest a relation with
patulum
but the female genital morphology of the specimen we analysed differs from the samples of
patulum
. Another entity has been described from the same area (roughly
5 km
south) as
Cochlostoma acutum
(
Caziot, 1908
)
. This is considered a synonym of
“
simrothi
” by
Boato
et al
. (1984)
but accepted as a species on its own by
Gargominy & Ripken (2006)
because found syntopically with
C.
(
T.
)
simrothi
. We did not analyse samples of this taxon (only seen in pictures). It cannot be excluded that it is not part of
Turritus
and instead belongs to the subgenus
Dalfreddia
Zallot
et al
., 2015
.