Cochlostoma Jan, 1830 revised: an overview of the subgenus Turritus Westerlund, 1883 and its species (Caenogastropoda, Cochlostomatidae) Author Zallot, Enrico 365E17AD-6938-4364-A526-F1BDA2663E10 Per Ligont 1, 33070 Budoia, Italy. ezallot@gmail.com Author Kamchev, Panche 9FA2BE01-3355-4340-B57C-9E2841D5E4C2 Naturalis Biodiversity Center, Darwinweg 2, 2333 CR Leiden, the Netherlands. panekamchev@gmail.com Author Schilthuizen, Menno 683D0AB7-CDD8-4FAA-94B6-436F3BFB8873 Naturalis Biodiversity Center, Darwinweg 2, 2333 CR Leiden, the Netherlands. & Taxon Expeditions, Rembrandtstraat 20, 2311 VW Leiden, the Netherlands. menno.schilthuizen@naturalis.nl Author Fehér, Zoltán E801EC76-8B1E-450B-993E-BBBE57C00EA9 WWF Hungary, Álmos vezér útja 69 / A, 1141 Budapest, Hungary. feher.zoltan.nhmus@gmail.com Author Mattia, Willy De C74A049D-9D7B-4A1F-878B-5BDFD104219F Central Research Laboratories, Burgring 7, 1010, Vienna, Austria; Department of Evolutionary Biology, University of Vienna, Djerassiplatz 1, 1030, Vienna, Austria. wdemattia@gmail.com Author Gittenberger, Edmund D786C279-FC92-4D08-AF16-F79A9705E0AE Naturalis Biodiversity Center, P. O. Box 9517, 2300 RA Leiden, the Netherlands. egittenberger@yahoo.com text European Journal of Taxonomy 2024 2024-03-21 927 1 163 https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2475/11013 journal article 292111 10.5852/ejt.2024.927.2475 4eb20c92-816d-4a89-bf2a-484791f2966d 2118-9773 10853688 B6E43365-FACA-49BA-8CCD-77E4BF8F0016 Cochlostoma (T.) agriotes (Westerlund in Paulucci, 1879 ) Figs 97 (yellow dots), 98G, 100–101 Pomatias ( patulus Drap. Subsp.) agriotes Westerlund in Paulucci, 1879: 20 . Types Not seen. Other specimens ITALY • 1- Mt Cucco ( 10 km south along the same mountain range, here assumed as topotypical ); 43.3646° N , 12.731° E ; 2010; Zallot leg.; EZ0001 • 2- Gualdo Tadino ; 43.2302° N , 12.7957° E ; 2013; Margelli leg.; EZ1139 • 9- Gole dell’Infernaccio ; 42.9397° N , 13.3159° E ; 2013; Margelli leg.; EZ1142 . Type locality ITALY • Marche , Pesaro and Urbino, Fonte Avellana . Westerlund reported as localities “Italia at Suellana and Saorgio”. Paulucci already noticed that “Suellana” is the misspelled name of the hermitage Fonte Avellana. “Saorgio”, which is the second locality cited by Westerlund was once in Italy, but it is now a French village in the Maritime Alps which is inhabited by another taxon (locus typicus of C. ( T. ) simrothi ) . Fig. 97. Distribution of samples in Clade B: green = Cochlostoma ( T. ) crosseanum ( Paulucci, 1879 ) ; yellow= C. ( T. ) agriotes ( Westerlund, 1879 ) ; red= C. ( T. ) affine (Benoit, 1876) . Black center for samples with amplified 16S. Fig. 98. Shell and female genitals of samples of clade B with amplified 16S. A . Cochlostoma ( T. ) crosseanum ( Paulucci, 1879 ) , Posta, via Salaria, I (EZ-0002). B . Gole del Velino 2, I (EZ-1014). C . Gole del Velino 1, I (EZ-1013). D . Roccalbegna, I (EZ-0980). E . Lucchio, I (WdM-6953). F . C. ( T. ) affine (Benoit, 1876) , Isnello, I (EZ-1016). G . C. ( T. ) agriotes ( Westerlund, 1879 ) , Monte Cucco, I (EZ-0001). Fig. 99. Cochlostoma ( T. ) affine (Benoit, 1876) , 1- Isnello, I (EZ-1016). Description SHELL . Closely spaced riblets on last part of protoconch. Teleoconch spotless with more or less strong, often whitish, irregularly spaced ribs. Whitish powder on whorls. Rather weak lip with gently inwardly bent columellar lobe covering almost completely umbilicus. Fig. 100. Cochlostoma ( T. ) agriotes ( Westerlund, 1879 ) , 1- Monte Cucco, I (EZ-0001). Fig. 101. Cochlostoma ( T. ) agriotes ( Westerlund, 1879 ) . A . 1- Monte Cucco, I (EZ-0001). B . 1- Monte Cucco, I (EZ-0001). C . 2- Gualdo Tadino, I (EZ-1139). D . 3- Gole dell’Infernaccio (EZ-1142). MEASUREMENTS . 29 ♀♀ : whorls= 7.1–9.2, H = 7.5–10.2 mm , H/W=2.54–3.04, roundness=0.1–0.18, ribs incl.= 52–64°, apert. incl. =15–24°, ribs/mm 1 st wh. =2–10, ribs/mm 4 th wh.=4–10. FEMALE GENITAL ORGANS . As in C. ( T .) affine . Remarks Cochlostoma ( T. ) agriotes is considered a subspecies of C. ( T. ) crosseanum in MolluscaBase (2023) but actually they have the same shell and female genital morphology. However, the Sicilian C. ( T. ) affine is equally similar and therefore, on morphological characters, it should be synonymized too. The p-distances between the samples of agriotes and crosseanum are very high (2.4–3.1%), higher than the one between the Sicilian affine and agriotes (1.7%). The assignment of populations to this species rather than to the nearby living C. ( T. ) crosseanum is purely tentative and based on geographical consideration.