Cochlostoma Jan, 1830 revised: an overview of the subgenus Turritus Westerlund, 1883 and its species (Caenogastropoda, Cochlostomatidae)
Author
Zallot, Enrico
365E17AD-6938-4364-A526-F1BDA2663E10
Per Ligont 1, 33070 Budoia, Italy.
ezallot@gmail.com
Author
Kamchev, Panche
9FA2BE01-3355-4340-B57C-9E2841D5E4C2
Naturalis Biodiversity Center, Darwinweg 2, 2333 CR Leiden, the Netherlands.
panekamchev@gmail.com
Author
Schilthuizen, Menno
683D0AB7-CDD8-4FAA-94B6-436F3BFB8873
Naturalis Biodiversity Center, Darwinweg 2, 2333 CR Leiden, the Netherlands. & Taxon Expeditions, Rembrandtstraat 20, 2311 VW Leiden, the Netherlands.
menno.schilthuizen@naturalis.nl
Author
Fehér, Zoltán
E801EC76-8B1E-450B-993E-BBBE57C00EA9
WWF Hungary, Álmos vezér útja 69 / A, 1141 Budapest, Hungary.
feher.zoltan.nhmus@gmail.com
Author
Mattia, Willy De
C74A049D-9D7B-4A1F-878B-5BDFD104219F
Central Research Laboratories, Burgring 7, 1010, Vienna, Austria; Department of Evolutionary Biology, University of Vienna, Djerassiplatz 1, 1030, Vienna, Austria.
wdemattia@gmail.com
Author
Gittenberger, Edmund
D786C279-FC92-4D08-AF16-F79A9705E0AE
Naturalis Biodiversity Center, P. O. Box 9517, 2300 RA Leiden, the Netherlands.
egittenberger@yahoo.com
text
European Journal of Taxonomy
2024
2024-03-21
927
1
163
https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2475/11013
journal article
10.5852/ejt.2024.927.2475
2118-9773
10853688
B6E43365-FACA-49BA-8CCD-77E4BF8F0016
Cochlostoma
(
T.
)
elegans
(
Clessin, 1879
)
Figs 72
(cyan dots), 73E, 77–81
Pomatias elegans
Clessin, 1879: 122–123
.
Pomatias
(
Eupomatias
)
elegans
var.
irrregularis
Wagner, 1897: 11
, pl. 1 fig. 11.
Pomatias
(
Eupomatias
)
elegans
var.
spectabilis
Wagner, 1897: 11
, pl. 1 fig. 12.
Pomatias
(
Eupomatias
)
elegans
var.
tumida
Wagner, 1897: 12
, pl. 1 fig. 13a–b.
Pomatias
(
Eupomatias
)
elegans
var.
similis
Wagner, 1897: 12
, pl. 1 fig. 14.
Pomatias
(
Eupomatias
)
elegans
var.
oostoma
Westerlund, 1883: 12
, pl. 1 fig. 10a–b.
Pomatias
(
Eupomatias
)
elegans
var.
imoschiensis
Wagner, 1906: 98–99
, pl. 3 fig. 1a–b.
Pomatias
(
Eupomatias
)
braueri
var.
latestriatum
Wagner, 1897: 10
, pl. 1 fig. 8.
Syntype
CROATIA
•
♀
; “
Cochlostoma
(
C.
)
elegans
(Clessin)
Orig. 1897
T.1 F.9–Kroatien:
Podgorje
,
Velebith
, Clessin 1889”;
SMF158949
.
Other specimens
CROATIA
•
syntype
of
C.
(
T.
)
elegans irregularis
;
Velebit
;
43.8634° N
,
15.83° E
;
A.J. Wagner
leg.;
SMF158959
/5
•
syntype
of
C.
(
T.
)
elegans tumida
; 10-
Visočica near Gospić
;
44.43° N
,
15.3713° E
;
A.J. Wagner
leg.;
SMF158970
/3
•
syntype
of
C.
(
T.
)
elegans imoschiensis
; 7-
Imoschi
,
Süddalmatien
;
43.4505° N
,
17.2091° E
;
Sturany
leg.;
SMF158972
/10
•
syntype
of
C.
(
T.
)
elegans irregularis
; 1?-
Velebit
;
44.5339° N
,
15.0772° E
;
A.J. Wagner
leg.;
NHMW25349
•
syntype
of
C.
(
T.
)
elegans imoschiensis
; 7-
Imotski
;
43.4473° N
,
17.214° E
; 1901;
Sturany
leg.;
NHMW34514
•
syntype
of
C.
(
T.
)
elegans tumida
; 8-
Rača
by
Sveta Jurai
;
44.9277° N
,
14.9199° E
;
A.J. Wagner
leg.;
NHMWK46907
•
syntype
of
C.
(
T.
)
braueri latestriatum
; 9-
VakanskiVrh
;
44.3661° N
,
15.509° E
;
A.J. Wagner
leg.;
NHMW66398
•
syntype
of
C.
(
T.
)
braueri spectabilis
; 11-
Lukovo Zugarje
;
44.26° N
,
15.12° E
;
A.J. Wagner
leg.;
MIZPASW
6590
•
2-
Senj
(topotypical);
44.9851° N
,
14.9311° E
; 1999;
De Mattia
leg.;
WdM1862
•
3-
Senj-Sveti Jurai
(topotypical);
44.9187° N
,
14.9241° E
; 2009;
De Mattia
leg.;
WdM6887
•
4-
Razbojište near Oltari
;
44.9084° N
,
14.9453° E
; 2009;
De Mattia
leg.;
WdM6896
•
5-
1.7 km
E of Vidovac
;
44.5246° N
,
15.0718° E
; 1999;
Fehér
and
Tamás
leg.;
HNHM91857
•
6-
Velika Paklenica
;
44.3326° N
,
15.4833° E
; 1999;
Fehér
and
Tamás
leg.;
HNHM91859
•
7-
Imotski
;
43.4505° N
,
17.2091° E
; 2008;
Dányi
,
Fehér
,
Kontschán
and
Murányi
leg.;
HNHM97265
.
Type locality
Clessin (1879)
wrote: “Diese Art wurde mir von Professor Stossich von Croatien mitgetheilt, ohne nähere Angabe des Fundortes. [This species was communicated to me by Professor Stossich of
Croatia
, without any further details of the place of discovery]”. The label
SMF-
158949 states: “
Kroatien
: Podgorje, Velebith, Clessin 1889”. This is a region at the foothill of the Velebit Mountains along the Croatian coast between Senj and Zrmanja, more than
130 km
long.
Description
SHELL
. Spaced riblets on last part of protoconch. Teleoconch spotless. Whitish not very prominent ribs often highly but irregularly inclined with respect to main axis. Mouth very inclined too, with moderately strong lip and flat columellar lobe leaving wide umbilicus well visible.
MEASUREMENTS
.
18 ♀♀
: whorls= 6.3–8.5, H =5.5–8.0 mm, H/W=2.01–2.57, roundness =0.09–0.17, ribs incl.= 45–58°, apert. incl. =18–33°, ribs/mm 1
st
wh. =3–17, ribs/mm 4
th
wh.=6–14.
FEMALE
GENITAL
ORGANS
. As in
C.
(
T.
)
nanum
.
Remarks
We synonymize all the so-called subspecies because they do not have their own exclusive ranges with diagnostic character states. We synonymize
C.
(
T.
)
braueri latestriatum
(
Wagner, 1897
)
with
C.
(
T.
)
elegans
despite the lack of anatomical and molecular data on the following ground: the
syntype
of
C.
(
T.
)
braueri latestriatum
from Vaganski Vrh (
Fig. 80A
) has a shell substantially differing from the other supposed
syntype
of “
latestriatum
” from Visočica peak (
Fig. 80B
. Vaganski Vrh and Visočica peak are mentioned as type localities in the original description). The latter presents the same shell morphology of the
“
dubium
” specimens from Mount Metla (
Fig. 89
). The same shell morphology and geographical proximity (roughly
20 km
along the same mountain range) between Visočica peak and Mount Metla induce to think that they are conspecific and both specimens of
C.
(
T.
)
dubium
(
Wagner, 1897
)
. Therefore, the
syntype
of this taxon is the specimen from Vaganski Vrh, but it is not clear why it has been defined as a subspecies of
“
braueri
” instead of
C.
(
T.
)
elegans
: comparing specimens as in
Fig. 81
, this appears much more probable than assigning it to a species living much further north and, as far as is known, restricted to a single mountain (Mount Klek). Moreover, Vakanski Vrh is only a few kilometers distant from Velika Paklenica, from where the sample is embedded in the phylogenetic tree in the
“
elegans
” branch.
Fig.77.
Types of subspecies of
Cochlostoma
(
T.
)
elegans
(
Clessin, 1879
)
at the Naturmuseum Senckenberg Frankfurt.
A
.
C.
(
T.
)
elegans elegans
, ♀, Podgorie, Velebith, HR (SMF-158949/4).
B
.
C.
(
T.
)
elegans imoschiensis
Wagner, 1906
, ♂, Imoschi, South Dalmatia, HR (SMF1-58972/10).
C
.
C.
(
T.
)
elegans irregularis
Wagner, 1897
, ♀, Velebith, HR (SMF-158959/5).
D
.
C.
(
T.
)
elegans oostoma
Westerlund, 1883
, ♂, Julian Alps (SMF-158961/2).
E
.
C.
(
T.
)
elegans tumida
Wagner, 1897
, probably ♀, Visočica in Velebith, Gospić, HR (SMF-158970/3).
Fig. 78.
Types of
Cochlostoma
(
T.
)
elegans
(
Clessin, 1879
)
.
A
.
C.
(
T.
)
elegans similis
Wagner, 1897
, ♂, Velebith? (MIZPAS no number).
B
.
C.
(
T.
)
elegans spectabilis
Wagner, 1897
, ♀, Lukovo, Zuganje, HR (MIZPAS-6590).
There are probably populations of
C.
(
T.
)
elegans
that have a subspecific status, but this has to be studied in more detail.