The Clavatulidae (Gastropoda, Conoidea) of the Miocene Paratethys Sea with considerations on fossil and extant Clavatulidae genera Author Harzhauser, Mathias 0000-0002-4471-6655 mathias.harzhauser@nhm-wien.ac.at Author Landau, Bernard 0000-0002-7768-8494 bernardmlandau@gmail.com Author Janssen, Ronald 0000-0002-4471-6655 mathias.harzhauser@nhm-wien.ac.at text Zootaxa 2022 2022-03-30 5123 1 1 172 http://dx.doi.org/10.11646/zootaxa.5123.1.1 journal article 10.11646/zootaxa.5123.1.1 1175-5326 6399827 036F6B4D-CDCC-4CD7-A914-9A1D8C7A097A ‘ Clavatula ’ santhai Kovács & Vicián, 2021 Figs 11G 1 –G 2 , 3N , 5 , 6 Clavatula santhai n. sp. — Kovács & Vicián 2021: 144 , pl. 3, figs 3–6. Type material. Holotype : HNHM , PAL 2021.20 .1, SL: 37.0 mm, MD : 13.5 mm , Letkés ( Hungary ), illustrated in Kovács & Vicián (2021 , pl. 3, figs 5–6), figs 11G 1 –G 2 , 3N . Revised description. Shell medium-sized, moderately slender fusiform with high spire; apical angle ~38°. Protoconch not preserved. Teleoconch of ten whorls. Earliest spire whorls with broad, smooth subsutural cord and row of suprasutural beads, separated by narrow spiral groove.Abapically, mid-portion widens and becomes concave, with axial riblets ending at the swollen suprasutural row of beads. Later whorls subcylindrical, subsutural collar weakly developed bearing small, pointed tubercles just below suture on last two whorls, weak central concavity with delicate spiral threads in some specimens, suprasutural beads strengthen to form axially elongated opisthocline tubercles, partly covered by succeeding whorl. Suture narrowly but deeply impressed, weakly undulating. Last whorl ~60% of the total height. Subsutural collar moderate in width, poorly delimited, with pointed tubercles adapically. Subsutural ramp, wide, weakly concave, delimited by tuberculose shoulder cord, convex below, weakly constricted at base. Siphonal fasciole moderately swollen, well delimited, twisted. Sculpture below ramp of seven primary cords. Shoulder cord tubercular, rest beaded. Peribasal cord slightly stronger, with secondary cords intercalated. Aperture moderately wide, ovate. Outer lip not thickened, smooth within. Siphonal canal moderately long, narrow. Anal canal moderately wide and deep, asymmetrically U-shaped, with apex mid-ramp. Columella excavated in upper third, straight below, slightly twisted at fasciole, smooth. Columellar and parietal callus thickened, sharply delimited, forming broad rim and narrow pseudumbilical chink. Discussion. This species is very similar to ‘ Clavatula’ camillae ( Hoernes & Auinger, 1891 ) and differs mainly in its more slender outline, less concave whorls, weaker spiral sculpture on the base, which is also less constricted, and lacks the double perifasciolar cords seen exclusively in ‘ C ’ camillae . Paleoenvironment. The occurrence at Letkés ( Hungary ) suggests a preference for inner neritic environments in the vicinity of corals ( Kovács & Vicián, 2013 ). Distribution in Central Paratethys. Badenian (middle Miocene): Pannonian Basin : Letkés ( Hungary ) ( Kovács & Vicián 2021 ).