The Beetle Family Carabidae of Costa Rica: Twenty­nine new species of Agra Fabricius 1801 (Coleoptera: Carabidae, Lebiini, Agrina) Author Erwin, Terry L. text Zootaxa 2002 119 1 68 journal article 10.5281/zenodo.162803 f514e964-2196-461c-84ff-64243643bc81 1175­5326 162803 Agra granodeoro Erwin , new species ( Figs. 19a, 19 b, 19c) . Male, COSTA RICA : Cartago, Turrialba, Chirripo, Grano de Oro , 1120 m , 09° 53 'N 83°38'W , September, (P. Campos)( INBio : CRI 000­935857). Diagnosis . This is the smallest member of the palmata species­group in Costa Rica and the only species with a narrow, tapered prothorax, black knees and tarsi. Description . Color and luster: Piceous. Legs rufotestaceous with black knees and tarsi. Antennomeres 1­2 and apex of 3­10 infuscated. Female unknown. Form: Head ( Fig. 19a ) behind eyes moderately long, sides tapered to abruptly obtuse angles. Pronotum widest at base, markedly narrowed just posterior to apical collar; disc sparsely punctate. Elytron with apex ( Fig. 19 b) markedly oblique, finely dentate suturally, sutural margin sinuate. Aedeagus ( Fig. 19 c) with two ostial membranes separated by a wide sclerotized bridge, endophallus without vestiture or armature; apex of phallus diamond­shaped. Size: moderately large, 18.1 mm in length, 4.4mm in width. Other specimens examined. The male Holotype is the only specimen I saw. Specific epithet. The specific epithet, granodeoro , is used as a noun in apposition, and refers in Spanish to a "golden seed," in reference to the brassy glint of the elytra and to the type locality. FIGURE 19. A. granodeoro , male, dorsal aspect: a) head; b) apex of elytron; c) aedeagus, dorsal (top), ventral (middle), left lateral (bottom) aspects. Measures (mm): HL = 3.3; WE = 2.2; AL = 5.0. Notes . The Holotype is from the Cordillera Volcánica Central Conservation Area. An illustration of palmata­group stylomere 2 is on the Agra atriperna web page.