Review and phylogeny of Cyrtomenus Amyot & Serville (Hemiptera: Cydnidae: Cydninae) based on morphological characters Author Avendaño, José Mauricio Laboratorio de Entomologia Sistemática. Departamento de Zoologia, Instituto de Biociências, Universidade Federal do Rio Grande do Sul (UFRGS), Brazil Author Correia, Maíra Andrade PPG Ecologia e Evolução, Instituto de Ciências Ambientais, Químicas e Farmacêuticas, Universidade Federal de São Paulo (UNIFESP), Brazil Author Grazia, Jocelia Laboratorio de Entomologia Sistemática. Departamento de Zoologia, Instituto de Biociências, Universidade Federal do Rio Grande do Sul (UFRGS), Brazil Author Schwertner, Cristiano Feldens Departamento de Ecologia e Biologia Evolutiva, Instituto de Ciências Ambientais, Químicas e Farmacêuticas, Universidade Federal de São Paulo (UNIFESP), Brazil & PPG Ecologia e Evolução, Instituto de Ciências Ambientais, Químicas e Farmacêuticas, Universidade Federal de São Paulo (UNIFESP), Brazil & Museu de Zoologia (MZUSP), Universidade de São Paulo, Brazil text Zootaxa 2024 2024-05-21 5453 3 301 340 http://dx.doi.org/10.11646/zootaxa.5453.3.1 journal article 296804 10.11646/zootaxa.5453.3.1 035c358a-0715-41a2-a6c9-27ebad4ea273 1175-5326 11234163 8B91667E-638D-4F5C-8A83-A806474228DF Cyrtomenus crassus Walker, 1867 ( Figs. 5E , 6L , 11 , 13 ) Cyrtomenus crassus Walker, 1867: 147 . Cyrtomenus obtusus Uhler, 1877: 369 ; (syn. by Froeschner, 1960 ). Cyrtomenus mirabilis ; Distant, 1880: 3; Signoret, 1881: 199; Uhler, 1886: 3; Van Duzee, 1917: 18; (syn. by Froeschner, 1960 ). Cyrtomenus castaneus ; Lethierry & Severin, 1893: 62; (syn. by Froeschner, 1960 ). Cyrtomenus vestigiatus Distant, 1903: 525 ; (syn. by Froeschner, 1960 ). Cyrtomenus (Cyrtomenus) crassus ; Froeschner, 1960: 533 . Material examined: USA : Arizona : 1♂ , 1♀ , Gila co. , Globe, D.K . Duncan , AMNH ; 1♀ , Santa Cruz co. , Nogales , 28-Jul-56, C. & M. Cazier , AMNH ; 1♂ , Santa Cruz co. , Patagonia. Sonoita River , 18-Jul-48, C. & P. Vaurie , 245268, AMNH ; MÉXICO : Chihuahua , 1♀ , Matachí , Matachí , 7-Jul-47, D. Rockefeller , 245246, AMNH ; Jalisco : 1♂ , Guadalajara , 5 Mi W La Venta , 4-Jul-53, C. & P. Vaurie , 245262, AMNH ; Nayarit : 1♂ , 1♀ , Compostela , 3 mi NW Las Varas , 11-Nov-50, R . Smith , AMNH ; 1♂ , Tepic , Tepic , 28-Jul-53, C. & P. Vaurie , AMNH ; 1♀ , Tepic , Tepic , 2-Aug-47 to 7-Aug-47, B. Malkin , AMNH ; Sinaloa , 1♂ , Mazatlán , Mazatlán , 2-Aug-53, C. & P. Vaurie , 246717, AMNH ; 1♂ , 1♀ , Mazatlán , Mazatlán , 22-Jul-54, M. Casier , AMNH ; Veracruz , 2♀ , San Andrés Tuxtla , Volcán San Martín , 29-May-51, I. Bassols , UNAM ; 1♀ , San Andrés Tuxtla , 26-May-51, A. Barrera , UNAM ; 1♂ , Córdoba , 4-Aug-65, A.B. Lau , NMNH ; CUBA : Soledad , 1♂ , 3-Jul-25, AMNH ; HONDURAS : El Paraiso : 1♂ , Danlí , 110 Km E Tegucigalpa. San Juan de Linaca , 6-Jul-83, Andrews , “Raiz de Zea maiz”, NMNH ; COLOMBIA : Bolívar , 1♀ , Zambrano , Hda Monterrey , 70 masl , 8-May-93, Fernández F. , IAvH-87270, IAVH ; 1♀ , Zambrano , Hda Monterrey , 70 masl , 12-Aug-93, Fernández F. , IAvH-05078, IAVH ; 1♀ , Zambrano , Hda Monterrey , 70 masl , 2- Sep-93, Fernández F. , IAvH-87165, IAVH ; 1♀ , Zambrano , Hda Monterrey , 70 masl , 9-Sep-93, Fernández F. , IAvH-87180, IAVH ; 1♂ , Zambrano , Hda Monterrey , 70 masl , 28-Oct-93, Fernández F. , IAvH-88119, IAVH ; Tolima : 1♀ , Mariquita , 30-Oct-05, R , Garzón , ICN ; 1♂ , Melgar , 11-Oct-95, F. Cárdenas , “en suelo”, ICN ; 1♂ , Melgar , 323 masl , 24-Apr-1976 , M. R . León , ICN . FIGURE 13 . C . crassus : (A) dorsal view; (B) head dorsal; (C) evaporatoria; (D) latero-ventral view. Scale bar: 1 mm. Diagnosis: Dorsal surface of the mandibular plates, smooth; Gonocoxites VIII longer than broad; and ovoid seminal receptacle of the spermatheca. Redescription. Total length: 7.6–9.0 mm. Head: surface noticeably convex ( Fig. 13A, B ), shining, with prominent rugae radiating from base of clypeus, minutely punctate, anterior outline rounded; mandibular plates longer than clypeus and nearly or quite contiguous in front of it, apices recurved; eyes projecting beyond sides of head by not more than one-third their width ( Fig. 13B ); bucculae less than half as high as labial II, latter without semicircular foliaceous lobe; labium apex reaching between meso- and metacoxae ( Fig. 13D ). Thorax: Pronotum with lateral submarginal row of 15 to 18 setigerous punctures; transverse impression weak, with medially interrupted, irregular row of coarse, close-set punctures; anterior lobe impunctate except for moderate punctures laterally and in subapical band, median subapical impression broad, very shallow; posterior lobe impunctate or with a few coarse punctures anteriorly ( Fig. 13A ). Scutellum disc with few to several widely scattered, coarse punctures. Hemelytron shining, uniformly punctate; clavus with one row of punctures; mesocorium with two rows of punctures paralleling claval suture; exocorium sparsely punctate; costa with six to ten setigerous punctures ( Fig. 13A, D ). Propleuron shining, with few distinct punctures in depression. Mesopleural evaporatorium not interrupted by pseudoperitreme; anterolateral area polished, impunctate, with few oblique rugae. Posterolateral area polished, impunctate ( Fig. 13C ). Legs densely covered by long setae; metatibia strongly dilated toward apex, greatest diameter there equal to that of protibia ( Fig. 13D ). Abdomen: Sternites III to VI polished, with rows of setigerous punctures across segments ( Fig. 13D ). Distribution: United States , Mexico , Cuba , Guatemala , Honduras , Costa Rica , Colombia First record— ( Fig. 11 ). Remarks: See remarks of C. ciliatus .