Macrodactylini (Coleoptera, Scarabaeidae, Melolonthinae): primary types of type species and taxonomic changes to the generic classification
Author
Fuhrmann, Juares
Author
Vaz-De-Mello, Fernando Z.
text
European Journal of Taxonomy
2017
2017-09-13
350
1
71
journal article
22059
10.5852/ejt.2017.350
52e4ae01-f51f-4c4a-8a14-9fb5b17788d6
2118-9773
3832682
8D14DBDE-AD13-445B-B2D0-221F19FC7C37
Faula
Blanchard, 1850
, revalidated
Faula
–
Lacordaire 1856: 253
(synonymy with
Ancistrosoma
). —
Geminger & Harold 1869: 1154
(synonymy with
Ceraspis
).
Type
species
Faula cornuta
Blanchard, 1850
(designation:
Evans 2003: 236
).
Diagnosis
Body usually brown or reddish-brown, opaque; pronotal anterior margin not beaded; pronotum−scutellum contact roundly sinuous (
Fig. 8D
); prosternum with an anterior longitudinal carina; protibia with 3–4 external teeth and without spur; meso- and metacoxae separate; metatarsomere V with internoproximal spine-like setae.
Remarks
The genera
Ancistrosoma
,
Chariodema
,
Faula
,
Manopus
,
Pectinosoma
and
Pseudopectinosoma
are similar to each other. They have a sinuous pronotum−scutellum contact and meso- and metacoxae separate.
Faula
is distinguished by the protibia with 3–4 external teeth (rarely with proximal tooth reduced) and without spur; male ventrite VI weakly prominent or flat and straight.
The present study revalidates
Faula
, formerly a synonym of
Ceraspis
, based on the following characters (opposition to
Ceraspis
): pronotum and elytron glabrous or sparsely setose, pronotum rarely densely setose, elytron of some species with longitudinal setose bands, at least in females (usually densely setose, sometimes setae covering the surface largely or entirely); pronotum−scutellum contact roundly sinuous (
Fig. 8D
) (acutely sinuous,
Fig. 8C
); protibia with 3–4 external teeth, some minor specimens with proximal tooth reduced (two teeth); meso- and metacoxae medially separate (contiguous).
Faula
comprises27species(13species from original combination and14species transferred from
Ceraspis
):
F
.
bicolor
(
Moser, 1919
)
comb. nov.
,
F
.
bivittata
(
Burmeister, 1855
)
comb. nov.
,
F
.
brunneipennis
Bates, 1887
,
F
.
centralis
Sharp, 1877
,
F
.
clypealis
(Frey, 1962)
comb. nov.
,
F
.
cornuta
Blanchard, 1850
,
F
.
hispida
Bates, 1887
,
F
.
immaculata
(
Burmeister, 1855
)
comb. nov.
,
F
.
innotata
Blanchard, 1850
,
F
.
insularis
Arrow, 1903
,
F
.
jaliscoensis
(Delgado & Navarette-Heredia, 2004)
comb. nov.
,
F
.
klenei
(Brenske, 1890)
comb. nov.
,
F
.
lineata
Waterhouse, 1879
,
F
.
macrophylla
(
Moser, 1919
)
comb. nov.
,
F
.
mexicana
Harold, 1863
,
F
.
modesta
(
Burmeister, 1855
)
comb. nov.
,
F
.
oaxacaensis
(Delgado, 2001)
comb. nov.
,
F
.
ohausi
(
Moser, 1921
)
comb. nov.
,
F
.
opacipennis
Moser, 1919
,
F
.
pilatei
Harold, 1863
,
F
.
quadrifoliata
(
Moser, 1919
)
comb. nov.
,
F
.
quadrimaculata
Blanchard, 1850
,
F
.
quadripustulata
Blanchard, 1850
,
F
.
rubiginosa
(Latreille, 1813)
comb. nov.
,
F
.
ruhli
(Brenske, 1890)
comb. nov.
,
F
.
rufoscutellata
(
Moser, 1919
)
comb. nov.
and
F
.
squamulifera
Moser, 1919
.