Macrodactylini (Coleoptera, Scarabaeidae, Melolonthinae): primary types of type species and taxonomic changes to the generic classification Author Fuhrmann, Juares Author Vaz-De-Mello, Fernando Z. text European Journal of Taxonomy 2017 2017-09-13 350 1 71 journal article 22059 10.5852/ejt.2017.350 52e4ae01-f51f-4c4a-8a14-9fb5b17788d6 2118-9773 3832682 8D14DBDE-AD13-445B-B2D0-221F19FC7C37 Schizochelus Blanchard, 1850 Gastrohoplus Moser, 1921: 165 , syn. nov . Type species: Gastrohoplus mirabilis Moser, 1921 (designation: monotypy). Type species Schizochelus flavescens Blanchard, 1850 (designation: Evans 2003: 346 ). Fig. 15. Schizochelus Blanchard, 1850 , ♂, habitus, lateral (without some appendages). A . Schizochelus flavescens Blanchard, 1850 . B . Schizochelus bicoloripes Blanchard, 1850 . C . Schizochelus mirabilis ( Moser, 1921 ) comb. nov. Scale bar = 1 mm. Fig. 16. Schizochelus Blanchard, 1850 . A–C . Male abdomen, lateroventral. D . Female abdomen, lateral. E–F . Aedeagus (lateral, parameres apex). G–L . Protibia−tarsus (male, female) (with detail of tarsus: I = dorsal view; K = ventral view). A , E−H . Schizochelus flavescens Blanchard, 1850 . B , I–J . Schizochelus bicoloripes Blanchard, 1850 . C–D , K–L . Schizochelus mirabilis ( Moser, 1921 ) comb. nov. Scale bars = 1 mm. Fig 17. Lectotypes (dorsal, labels). A–B . Agaocnemis pruina Moser, 1918 . C–D . Corminus canescens Burmeister, 1855 . E–F . Anomalonyx uruguayensis Moser, 1921 . G–H . Barybas nana Blanchard, 1850 . I–J . Ctilocephala pelluscens Burmeister, 1855 . K–L . Pseudohercitis viridiaenea Moser, 1921 . M–N . Barybas volvulus Burmeister, 1855 . O–P . Calodactylus tibialis Blanchard, 1850 . Scale bars = 2 mm. Fig 18. Lectotypes (dorsal, labels). A–B . Ceraspis pruinosa LePeletier de Saint-Fargeau & Audinet- Serville, 1828. C–D . Ceratolontha venezuelae Arrow, 1948 . E–F . Chariodactylus chacoensis Moser, 1919 . G–H . Philochlaenia virescens Blanchard, 1842 . I–J . Clavipalpus dejeani Laporte, 1832 . K–L . Ctenotis obesa Burmeister, 1855 . M–N . Euryaspis gaudichaudii Blanchard, 1850 . O–P . Faula cornuta Blanchard, 1850 . Scale bars = 2 mm. Fig 19. Lectotypes (dorsal, labels). A–B . Mallotarsus spadiceus Blanchard, 1850 . C–D . Manodactylus gaujoni Moser, 1919 . E–F . Manopus biguttatus Conte de Castelnau, 1840 . G–H . Oedichira pachydactyla Burmeister, 1855 . I–J . Amphicrania ursina Burmeister, 1855 . K–L . Pectinosoma elongata Arrow, 1913 . M–N . Aulanota sulcipennis Moser, 1924 . O–P . Melolontha rufipennis Fabricius, 1801 . Scale bars = 2 mm. Fig 20. Lectotypes (dorsal, labels). A–B . Pachycerus castaneipennis Guérin-Méneville, 1831 . C–D . Anomalochilus singularis Blanchard, 1850 . E–F . Demodema fallax Blanchard, 1850 . G–H . Plectris tomentosa LePeletier de Saint-Fargeau & Audinet-Serville, 1828 . I–J . Gama grandicornis Blanchard, 1850 . K–L . Pachylotoma viridis Blanchard, 1850 . M–N . Serica marmorea Guérin-Méneville, 1831 . O–P . Rhinaspoides aeneofusca Moser, 1919 . Scale bars = 2 mm. Fig 21. Primary types. A–F . Lectotypes (dorsal, labels). A–B . Ulomenes hypocrita Blanchard, 1850 . C–D . Gastrohoplus mirabilis Moser, 1921 . E–F . Schizochelus flavescens Blanchard, 1850 . G–K . Syntype of Hercitis pygmaea Burmeister, 1855 (by Holger Dombrow). G . Dorsal. H . Lateral. I . Frontal. J . Posterior. K . Labels. Scale bars: A, C, E = 2 mm; G–J without scale (specimen about 4–4.2 mm according to original description). Diagnosis Clypeal anteroventral area strongly reduced ( Fig. 15 , similar to Fig. 11H ); pronotal anterior and posterior margins not beaded; prosternum with an anterior longitudinal carina; protibia with 1–2 external teeth (male with tooth II absent or reduced) and with a spur ( Fig. 16 G–L); male metatibia without spur; male ventrites I–V longitudinally concave ( Fig. 16 A–C); female abdomen sinuous in lateral view ( Fig. 16D ); tarsal claws with a proximal tooth ( Fig. 16 G–L). Remarks The genera Isonychus and Schizochelus are similar to each other. Schizochelus is distinguished as follows (opposition to Isonychus ): male metatibial spur absent (metatibia with 1–2 spurs); male ventrites I–V with medial concavity ( Fig. 16 A–C) (without), male ventrite VI straight and with short and sparse setae (straight or laterally enlarged, and with or without lateral comb of long setae); female abdomen sinuous in lateral view ( Fig. 16D ) (convex). The present study proposes the synonymy of Gastrohoplus with Schizochelus , see chapter on “Male morphology and nomenclature” (below) for more information about the new synonymy.