<document id="82EBA698176C29497E2F5A8AB9ECFC05" ID-DOI="10.5852/ejt.2024.948.2637" ID-ISSN="2118-9773" ID-Zenodo-Dep="13511901" ID-ZooBank="9985FD07-5280-41D9-B982-B175085AE5F8" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1724950631476" checkinUser="plazi" docAuthor="Gautam, R. K., Uniyal, V. P. &amp; Wood, Thomas J." docDate="2024" docId="03DE87B4FF87FFB70EDE6850FD12FA54" docLanguage="en" docName="EJT.2024.948.1-59.pdf" docOrigin="European Journal of Taxonomy 948" docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2637/12143" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Andrena leaena Cameron 1907" docType="treatment" docVersion="2" lastPageNumber="43" masterDocId="FFE7FFCCFFADFF9D0F396A02FF8EFF80" masterDocTitle="A critical revision of the Andrena Fabricius, 1775 of India, with the description of two new species (Hymenoptera: Andrenidae) from Uttarakhand" masterLastPageNumber="59" masterPageNumber="1" pageNumber="43" updateTime="1724951248699" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="DF204B9B135064B0F985CDD3D018A346">A critical revision of the Andrena Fabricius, 1775 of India, with the description of two new species (Hymenoptera: Andrenidae) from Uttarakhand</mods:title>
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<mods:namePart id="C19906126A2DD111FF498CC0DA1B91B5">Gautam, R. K.</mods:namePart>
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<mods:namePart id="747E571CF465F3721862A903E604DC25">Wood, Thomas J.</mods:namePart>
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<paragraph id="8BC836A2FF87FFB70EDE6850FBC3FDEC" blockId="42.[487,1101,593,620]" box="[487,1101,593,620]" pageId="42" pageNumber="43">
<emphasis id="B903EAB0FF87FFB70EDE6850FD82FDEC" bold="true" box="[487,524,594,620]" pageId="42" pageNumber="43">34.</emphasis>
<taxonomicName id="4C774D21FF87FFB70D2A6853FBC3FDEC" ID-CoL="DS3H" authority="Cameron, 1907" authorityName="Cameron" authorityYear="1907" box="[531,1101,593,620]" class="Insecta" family="Andrenidae" genus="Andrena" isHigherUncertain="true" kingdom="Animalia" order="Hymenoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="leaena">
<emphasis id="B903EAB0FF87FFB70D2A6853FDF0FDEC" bold="true" box="[531,638,593,620]" italics="true" pageId="42" pageNumber="43">Andrena</emphasis>
(incertae sedis) 
<emphasis id="B903EAB0FF87FFB70C796853FC1FFDEC" bold="true" box="[832,913,593,620]" italics="true" pageId="42" pageNumber="43">leaena</emphasis>
<bibRefCitation id="EFE64B53FF87FFB70CA16850FBC3FDEC" author="Cameron P." box="[920,1101,594,620]" pageId="42" pageNumber="43" pagination="1001 - 1012" refId="ref27543" refString="Cameron P. 1907. Description of a new genus and some new species of Hymenoptera captured by Lieut. - Col. C. G. Nurse at Deesa, Matheran and Ferozepore. Journal of Bombay Natural History Society 17: 1001 - 1012." type="journal article" year="1907">Cameron, 1907</bibRefCitation>
</taxonomicName>
</paragraph>
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<subSubSection id="C36D6529FF87FFB70F846891FEBDFD6F" pageId="42" pageNumber="43" type="reference_group">
<paragraph id="8BC836A2FF87FFB70F846891FBECFD2D" blockId="42.[189,1122,659,686]" box="[189,1122,659,686]" pageId="42" pageNumber="43">
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<treatmentCitation id="0AD610B3FF87FFB70F846891FD0FFD2E" author="Cameron P." box="[189,641,659,686]" page="1002" pageId="42" pageNumber="43" year="1907">
<taxonomicName id="4C774D21FF87FFB70F846891FD0FFD2E" ID-CoL="DS3H" authority="Cameron, 1907: 1002" authorityName="Cameron, 1907:" authorityPageNumber="1002" authorityYear="1002" box="[189,641,659,686]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="leaena">
<emphasis id="B903EAB0FF87FFB70F846891FEF6FD2D" box="[189,376,659,685]" italics="true" pageId="42" pageNumber="43">Andrena leaena</emphasis>
<bibRefCitation id="EFE64B53FF87FFB70EB96896FD0FFD2E" author="Cameron P." box="[384,641,660,686]" pageId="42" pageNumber="43" pagination="1001 - 1012" refId="ref27543" refString="Cameron P. 1907. Description of a new genus and some new species of Hymenoptera captured by Lieut. - Col. C. G. Nurse at Deesa, Matheran and Ferozepore. Journal of Bombay Natural History Society 17: 1001 - 1012." type="journal article" year="1907">Cameron, 1907: 1002</bibRefCitation>
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, 
<specimenCount id="9D71FD2BFF87FFB70DB46896FD2DFD2E" box="[653,675,660,686]" count="1" pageId="42" pageNumber="43" type="female">♀</specimenCount>
[
<collectingCountry id="F3607632FF87FFB70D8B6891FD7BFD2D" box="[690,757,659,685]" name="India" pageId="42" pageNumber="43">India</collectingCountry>
: Punjab, NHMUK, examined].
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BC836A2FF87FFB70F8468D7FEBDFD6F" blockId="42.[189,307,725,751]" box="[189,307,725,751]" pageId="42" pageNumber="43">
<heading id="D08081CEFF87FFB70F8468D7FEBDFD6F" bold="true" box="[189,307,725,751]" fontSize="11" level="3" pageId="42" pageNumber="43" reason="3">
<emphasis id="B903EAB0FF87FFB70F8468D7FEBDFD6F" bold="true" box="[189,307,725,751]" pageId="42" pageNumber="43">Listed by</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C36D6529FF87FFB70F846906FDEFFCC1" pageId="42" pageNumber="43" type="description">
<paragraph id="8BC836A2FF87FFB70F846906FDEFFCC1" blockId="42.[189,1399,771,833]" pageId="42" pageNumber="43">
<bibRefCitation id="EFE64B53FF87FFB70F846906FE0DFC9E" author="Cameron P." box="[189,387,772,798]" pageId="42" pageNumber="43" pagination="1001 - 1012" refId="ref27543" refString="Cameron P. 1907. Description of a new genus and some new species of Hymenoptera captured by Lieut. - Col. C. G. Nurse at Deesa, Matheran and Ferozepore. Journal of Bombay Natural History Society 17: 1001 - 1012." type="journal article" year="1907">Cameron (1907)</bibRefCitation>
; 
<bibRefCitation id="EFE64B53FF87FFB70EA86906FDA5FC9D" author="Batra S. W. T." box="[401,555,771,798]" pageId="42" pageNumber="43" pagination="164 - 177" refId="ref27165" refString="Batra S. W. T. 1967. Crop pollination and the flower relationships of the wild bees of Ludhiana, India. Journal of the Kansas Entomological Society 40: 164 - 177." type="journal article" year="1967">Batra (1967)</bibRefCitation>
; 
<bibRefCitation id="EFE64B53FF87FFB70D006901FC23FC9E" author="Tadauchi O. &amp; Matsumura T." box="[569,941,771,798]" pageId="42" pageNumber="43" pagination="1 - 20" refId="ref30285" refString="Tadauchi O. &amp; Matsumura T. 2007. The genus Andrena collected from Nepal (Hymenoptera, Andrenidae) with redescriptions of some types of Andrena described from North India. Esakia 47: 1 - 20. https: // doi. org / 10.5109 / 8322" type="journal article" year="2007">Tadauchi &amp; Matsumura (2007)</bibRefCitation>
; 
<bibRefCitation id="EFE64B53FF87FFB70C826901FB31FC9E" author="Chandra K. &amp; Saint J. &amp; Gupta D." box="[955,1215,771,798]" pageId="42" pageNumber="43" pagination="153 - 165" refId="ref27672" refString="Chandra K., Saint J. &amp; Gupta D. 2019. Insecta: Hymenoptera: Apoidea (Bees). Fauna of Punjab, State Fauna Series 23: 153 - 165." type="book chapter" year="2019">
Chandra 
<emphasis id="B903EAB0FF87FFB70B106906FBEAFC9D" box="[1065,1124,771,797]" italics="true" pageId="42" pageNumber="43">et al.</emphasis>
(2019)
</bibRefCitation>
; 
<bibRefCitation id="EFE64B53FF87FFB70BF46906FE9FFCC1" author="Meena L. K. &amp; Dey D." pageId="42" pageNumber="43" pagination="443 - 452" refId="ref29114" refString="Meena L. K. &amp; Dey D. 2019. Annotated checklist of non- Apis bees of the genus Andrena (Hymenoptera: Andrenidae) from India. Indian Journal of Entomology 81: 443 - 452 https: // doi. org / 10.5958 / 0974 - 8172.2019.00095.6" type="journal article" year="2019">Meena &amp; Dey (2019)</bibRefCitation>
; 
<bibRefCitation id="EFE64B53FF87FFB70E226925FDD3FCC1" author="Ascher J. &amp; Pickering J." box="[283,605,807,833]" pageId="42" pageNumber="43" refId="ref26955" refString="Ascher J. &amp; Pickering J. 2023. Discover Life bee Species Guide and World Checklist (Hymenoptera: Apidae: Anthophila). Availbable from https: // www. discoverlife. org / mp / 20 q? guide = Apoidea _ species &amp; flags = HAS [accessed 25 Jul. 2024]." type="url" year="2023">Ascher &amp; Pickering (2023)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C36D6529FF87FFB70F84696AFEFEFBE7" pageId="42" pageNumber="43" type="materials_examined">
<paragraph id="8BC836A2FF87FFB70F84696AFEE3FC02" blockId="42.[189,365,872,898]" box="[189,365,872,898]" pageId="42" pageNumber="43">
<heading id="D08081CEFF87FFB70F84696AFEE3FC02" bold="true" box="[189,365,872,898]" fontSize="11" level="3" pageId="42" pageNumber="43" reason="3">
<emphasis id="B903EAB0FF87FFB70F84696AFEE3FC02" bold="true" box="[189,365,872,898]" pageId="42" pageNumber="43">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="8BC836A2FF87FFB70FD36994FEDEFC30" blockId="42.[189,1152,918,980]" box="[234,336,918,944]" pageId="42" pageNumber="43">
<emphasis id="B903EAB0FF87FFB70FD36994FEDEFC30" bold="true" box="[234,336,918,944]" pageId="42" pageNumber="43">
<typeStatus id="54CC8800FF87FFB70FD36994FEDEFC30" box="[234,336,918,944]" pageId="42" pageNumber="43" type="syntype">Syntype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8BC836A2FF87FFB70F8469B8FB0EFC54" blockId="42.[189,1152,918,980]" box="[189,1152,954,980]" pageId="42" pageNumber="43">
<materialsCitation id="3B1F3CFFFF87FFB70F8469B8FBF2FC54" box="[189,1148,954,980]" collectingDate="1898-03-01" collectingDateMax="1898-03-31" collectingDateMin="1898-03-01" collectionCode="NHMUK" collectorName="C. G. Nurse" country="India" location="INDIA" pageId="42" pageNumber="43" specimenCount="1" specimenCount-female="1" typeStatus="syntype">
<collectingCountry id="F3607632FF87FFB70F8469B8FE9EFC54" box="[189,272,954,980]" name="India" pageId="42" pageNumber="43">INDIA</collectingCountry>
<emphasis id="B903EAB0FF87FFB70E2169B8FEACFC54" bold="true" box="[280,290,954,980]" pageId="42" pageNumber="43">•</emphasis>
<specimenCount id="9D71FD2BFF87FFB70E1369B8FED9FC54" box="[298,343,954,980]" count="1" pageId="42" pageNumber="43" type="female">1 ♀</specimenCount>
; Ferozepore [Firozpur]; 
<date id="FFC91062FF87FFB70D4169B8FCB8FC54" box="[632,822,954,980]" pageId="42" pageNumber="43" value="1898-03-01" valueMax="1898-03-31" valueMin="1898-03-01">
<collectingDate id="EF8DE98AFF87FFB70D4169B8FCB8FC54" box="[632,822,954,980]" pageId="42" pageNumber="43" value="1898-03-01" valueMax="1898-03-31" valueMin="1898-03-01">1–31 Mar. 1898</collectingDate>
</date>
; 
<collectorName id="26825374FF87FFB70C7B69B8FC49FC54" box="[834,967,954,980]" pageId="42" pageNumber="43">C.G. Nurse</collectorName>
leg.; 
<collectionCode id="ED66AE67FF87FFB70B3069B8FBF2FC54" box="[1033,1148,954,980]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="42" pageNumber="43" type="Museum">NHMUK</collectionCode>
</materialsCitation>
.
</paragraph>
<paragraph id="8BC836A2FF87FFB70F8469F9FE72FB95" blockId="42.[189,508,1019,1045]" box="[189,508,1019,1045]" pageId="42" pageNumber="43">
<heading id="D08081CEFF87FFB70F8469F9FE72FB95" bold="true" box="[189,508,1019,1045]" fontSize="11" level="3" pageId="42" pageNumber="43" reason="3">
<emphasis id="B903EAB0FF87FFB70F8469F9FE72FB95" bold="true" box="[189,508,1019,1045]" pageId="42" pageNumber="43">Other material examined</emphasis>
</heading>
</paragraph>
<paragraph id="8BC836A2FF87FFB70F846E28FEFEFBE7" blockId="42.[189,1399,1066,1127]" pageId="42" pageNumber="43">
<materialsCitation id="3B1F3CFFFF87FFB70F846E28FEE2FBE7" collectingDate="1898-03-01" collectingDateMax="1898-03-31" collectingDateMin="1898-03-01" collectionCode="NHMUK" collectorName="C. G. Nurse" country="India" location="INDIA" pageId="42" pageNumber="43" specimenCount="1" specimenCount-female="1">
<collectingCountry id="F3607632FF87FFB70F846E28FE80FBC4" box="[189,270,1066,1092]" name="India" pageId="42" pageNumber="43">INDIA</collectingCountry>
<emphasis id="B903EAB0FF87FFB70E266E28FEA7FBC4" bold="true" box="[287,297,1066,1092]" pageId="42" pageNumber="43">•</emphasis>
<specimenCount id="9D71FD2BFF87FFB70E006E28FEE1FBC4" box="[313,367,1066,1092]" count="1" pageId="42" pageNumber="43" type="female">1 ♀</specimenCount>
; Ferozepore [Firozpur]; 
<date id="FFC91062FF87FFB70D956E28FCF5FBC4" box="[684,891,1066,1092]" pageId="42" pageNumber="43" value="1898-03-01" valueMax="1898-03-31" valueMin="1898-03-01">
<collectingDate id="EF8DE98AFF87FFB70D956E28FCF5FBC4" box="[684,891,1066,1092]" pageId="42" pageNumber="43" value="1898-03-01" valueMax="1898-03-31" valueMin="1898-03-01">1–31 Mar. 1898</collectingDate>
</date>
; 
<collectorName id="26825374FF87FFB70CA96E28FB90FBC4" box="[912,1054,1066,1092]" pageId="42" pageNumber="43">C.G. Nurse</collectorName>
leg.; 
<collectionCode id="ED66AE67FF87FFB70B4B6E28FB6FFBC4" box="[1138,1249,1066,1092]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="42" pageNumber="43" type="Museum">NHMUK</collectionCode>
(previously undetermined)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C36D6529FF87FFB70F846E8DFC62FAE4" pageId="42" pageNumber="43" type="discussion">
<paragraph id="8BC836A2FF87FFB70F846E8DFEBEFB29" blockId="42.[189,304,1167,1193]" box="[189,304,1167,1193]" pageId="42" pageNumber="43">
<heading id="D08081CEFF87FFB70F846E8DFEBEFB29" bold="true" box="[189,304,1167,1193]" fontSize="11" level="3" pageId="42" pageNumber="43" reason="3">
<emphasis id="B903EAB0FF87FFB70F846E8DFEBEFB29" bold="true" box="[189,304,1167,1193]" pageId="42" pageNumber="43">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8BC836A2FF87FFB70F846EBFFC62FAE4" blockId="42.[189,1400,1213,1380]" pageId="42" pageNumber="43">
Although the type of 
<taxonomicName id="4C774D21FF87FFB70EFB6EBFFDBBFB57" authorityName="Cameron, 1907:" authorityYear="1002" box="[450,565,1213,1239]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="leaena">
<emphasis id="B903EAB0FF87FFB70EFB6EBFFDBBFB57" box="[450,565,1213,1239]" italics="true" pageId="42" pageNumber="43">A. leaena</emphasis>
</taxonomicName>
is conserved, its subgeneric affiliation remains unclear, and the male has never been described. Additional study is required to clarify the nature of this species. Though the specimen in the NHMUK type collection is labelled “type”, it is not automatically the 
<typeStatus id="54CC8800FF87FFB70BFB6F01FAA7FA9D" box="[1218,1321,1283,1309]" pageId="42" pageNumber="43" type="holotype">holotype</typeStatus>
and is only a 
<typeStatus id="54CC8800FF87FFB70E2D6F25FEE1FAC1" box="[276,367,1319,1345]" pageId="42" pageNumber="43" type="syntype">syntype</typeStatus>
as 
<bibRefCitation id="EFE64B53FF87FFB70EA46F25FDEFFAC1" author="Cameron P." box="[413,609,1319,1345]" pageId="42" pageNumber="43" pagination="1001 - 1012" refId="ref27543" refString="Cameron P. 1907. Description of a new genus and some new species of Hymenoptera captured by Lieut. - Col. C. G. Nurse at Deesa, Matheran and Ferozepore. Journal of Bombay Natural History Society 17: 1001 - 1012." type="journal article" year="1907">Cameron (1907)</bibRefCitation>
indicates that specimens come from both Ferozepore and Deesa, indicating that multiple individuals were used to describe the species.
</paragraph>
</subSubSection>
<subSubSection id="C36D6529FF87FFB70F846F89FD12FA54" box="[189,668,1419,1492]" pageId="42" pageNumber="43" type="distribution">
<paragraph id="8BC836A2FF87FFB70F846F89FED6FA25" blockId="42.[189,344,1419,1445]" box="[189,344,1419,1445]" pageId="42" pageNumber="43">
<heading id="D08081CEFF87FFB70F846F89FED6FA25" bold="true" box="[189,344,1419,1445]" fontSize="11" level="3" pageId="42" pageNumber="43" reason="3">
<emphasis id="B903EAB0FF87FFB70F846F89FED6FA25" bold="true" box="[189,344,1419,1445]" pageId="42" pageNumber="43">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BC836A2FF87FFB70F846FB8FD12FA54" blockId="42.[189,668,1466,1492]" box="[189,668,1466,1492]" pageId="42" pageNumber="43">
<collectingCountry id="F3607632FF87FFB70F846FB8FF77FA54" box="[189,249,1466,1492]" name="India" pageId="42" pageNumber="43">India</collectingCountry>
(Punjab, 
<collectingRegion id="49B3F840FF87FFB70E526FB8FE4BFA54" box="[363,453,1466,1492]" country="India" name="Gujarat" pageId="42" pageNumber="43">Gujarat</collectingRegion>
) (
<bibRefCitation id="EFE64B53FF87FFB70EE26FB8FD01FA54" author="Cameron P." box="[475,655,1466,1492]" pageId="42" pageNumber="43" pagination="1001 - 1012" refId="ref27543" refString="Cameron P. 1907. Description of a new genus and some new species of Hymenoptera captured by Lieut. - Col. C. G. Nurse at Deesa, Matheran and Ferozepore. Journal of Bombay Natural History Society 17: 1001 - 1012." type="journal article" year="1907">Cameron 1907</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>