establishment of five new genera from Australia, New Caledonia and New Zealand Author Gibbs, George W. text Zootaxa 2010 2520 1 48 journal article 10.5281/zenodo.196244 aa02f8e0-115c-4e5e-8e37-5912550d8dcf 1175-5326 196244 Tasmantrix tasmaniensis Gibbs , sp. nov. ( Figs 1 F, 12A–E, 20) Type material. Holotype : ɗ, Australia : 16 km NE Strahan, Tasmania, 42°09´S 145°28´E , 240 m , 23 Dec 1962 , I.F.B. Common ( ANIC ). Paratypes : 3ɗ 2 km SSW Renison Bell, Tasmania, 41°49´S 145°24´E , 260 m , 14 Dec 1981 , G.W. Gibbs; 3ɗ 7 km NE Zeehan, Tasmania, 41°51´S 145°25´E , 22 Nov 1988 , P.B.McQuillan & E.S. Nielsen; 1ɗ Strahan, Tasmania, 5 Feb 1989 , I.F.B. Common. (all specimens ANIC ) Material examined. Type series only. Female unknown. Diagnosis. Unique maculation: dark coppery iridescent ground colour with four white blotches, two on costa, two toward dorsum ( Fig 1 F). Male sclerite 9 forming a complete ring ( Figs. 12 A & B); valvae tapered with a minute mucron on inner apex ( Fig. 12 B); tergum 10 tapering to blunt apex ( Fig. 12 A). Description. Head interocular index of male 0.8. Male antenna with 52 (51–52) flagellomeres, scape and pedicel vestiture largely of lamellar scales, bronzy brown, a few longer creamy-yellow piliform scales overlaying the lamellar scales. First 3 flagellomeres filiform with vestiture of black lamellar scales, remainder of extreme moniliform, black; each ascoid with 7 curved branches (at mid-length of antenna). Mandibles functional, maxillary palps moderate length, 1.3x head width at compound eyes. Head capsule brown with dense tufts of creamy-white piliform scales. Lamellar scales of palps silvery white. Tegulae with shining bronzy-brown lamellar scales. Dorsum of thorax with shining bronzy-black lamellar scales; scales of coxae, femora and tibiae largely shining white, but fore-tibia and hind-tibia with bronzy-grey scales along anterior surface; fore-tarsus bronzy-grey, mid-tarsus grey with indistinct white bands, hind-tarsus dark bronzy-grey. Abdomen silvery-grey scaled. Forewing length of male 3.7mm (3.5–3.9). Maculation unique ( Fig. 1 F); ground colour rich brown with strong coppery-bronze iridescence. Basal costal streak absent but with four primary shining white fasciae and two secondary patches; a large oblique costal blotch at mid-length, separated from a smaller blotch on the termen in the same position; two similar blotches at three quarters on costa and termen, but with rows of black scales along the veins; a small, indistinct streak of paler scales about halfway along the cubital sulcis and a few pale scales in apex of wing. Fringes grey-brown with white tips but wholly white adjacent to the fasciae. Hindwing with 3 frenular bristles; dark grey scaled with bronzy lustre. Male abdomen and genitalia. [G920] ( Figs 12 A–E) No trace of a transverse sclerite on dorsum of A1. Exit area of S5 gland developed into a typical sabatincoid protuberance bearing about 17 long piliform scales. S8 not evident as a discrete sclerite although possibly represented by a small flange at the anterodorsal corner of sclerite 9, extending beyond the thickened margin and bearing a few microsetae. Sclerite 9 relatively large, 1.5x length of S6 along ventral mid-line, developed as a complete ring, with a broad band across the dorsum, unique in this genus; anterior concave margin thickened. Valvae simple, tapering, 2.2x longer than wide, with a small incurved mucron at the apex; inner surface uniformly clothed with curved setae, those at apex strongly orientated as ‘retro-setae’. Tergum 10 simple, hood-like, tapering to a blunt apex, a little short of the valve length; a small area of short retro-setae present on either side of the apex, on the ventral aspect. Anal cone well developed with lateral sclerite bearing microtrichia. Phallus average for the calliplaca -group, 2.2x length of S6; gonopore terminal with a gaping ‘fish-mouthed’ aperture surrounded by radial folds, a reduced ventral bulb present; phallocrypt densely clothed with acutely pointed scales. FIGURE 12. Male genitalia of Tasmantrix tasmaniensis sp. nov. : A, dorsal; B, lateral; C, ventral; D, phallus, lateral; E, distal apex of phallus enlarged; and Tasmantrix thula sp. nov : F, dorsal; G, lateral; H, ventral. Insets show specialised brush setae. Remarks. This, the largest of the calliplaca -group of species, is also the most straightforward to identify. Interestingly, it is the only Tasmantrix species in which the male exhibits a sabatincoid type of S5 gland protuberance. It occurs associated with Nothofagus cunninghamii forest remnants. Etymology. Species name derived from its geographic location in Tasmania. Distribution ( Fig. 20 ). Australia , in wet forests of western Tasmania, 41°49´S to 42°09´S , between 22 November and 5 February .