A review of the genus Sphingius Thorell, 1890 from India (Araneae: Liocranidae) Author Sankaran, Pradeep M. Division of Arachnology, Department of Zoology, Sacred Heart College, Thevara, Cochin, Kerala 682 013, India Author Caleb, John T. D. 0000-0002-9471-9467 Zoological Survey of India, Prani Vigyan Bhawan, M-Block, New Alipore, Kolkata, West Bengal 700 053, India caleb 87 woodgate @ gmail. com; https: // orcid. org / 0000 - 0002 - 9471 - 9467 caleb87woodgate@gmail.com Author Sebastian, Pothalil A. 0000-0002-4936-4310 Division of Arachnology, Department of Zoology, Sacred Heart College, Thevara, Cochin, Kerala 682 013, India & drpothalil @ rediffmail. com; https: // orcid. org / 0000 - 0002 - 4936 - 4310 drpothalil@rediffmail.com text Zootaxa 2020 2020-12-23 4896 4 505 522 journal article 9199 10.11646/zootaxa.4896.4.3 6592cf39-be44-44f4-842d-e0126d8640ad 1175-5326 4387671 0824AFA4-4E8B-419B-972C-0FA0A88538FF Inthaeron longipes ( Gravely, 1931 ) comb. nov. Figs 10–11 FIGURE 8. Sphingius sp.. A, C Ƌ: A habitus, dorsal; C eye group, dorsal. B, D ♀: B habitus, dorsal; D eye group, dorsal. E label from specimen bottle. F illustrations of ‘ S. caniceps ’ adopted from Gravely (1931) . Scale bars: A, 1 mm; B, 2 mm; C, 0.2 mm; D, 0.5 mm. Sphingius longipes Gravely, 1931: 270 , fig. 19A–B; Majumder & Tikader, 1991: 148 , figs 311–314. Type material. Syntype Ƌ from INDIA : Kerala : Palakkad: Pattambi (formerly in Malabar district ): beside the traveller’s bungalow, leg. F.H. Gravely , 24–28 May 1930 , deposited in NZC-ZSI (no register number), examined ( Fig. 10E ) . Justification of the transfer. Detailed examination of the syntype of S. longipes Gravely, 1931 revealed that it has the typical somatic morphology for Cithaeronidae Simon, 1893 ( Fig. 10 A–D) and the diagnostic features of the monotypic cithaeronid genus Inthaeron Platnick, 1991 , as illustrated for Inthaeron rossi Platnick, 1991 , the type species of the genus: highly coiled embolus, and embolus with a fringed flange (compare Fig. 11 A–C with Platnick & Gajbe 1994 : figs 1–3). Based on these observations, we propose to transfer S. longipes to Inthaeron . Diagnosis. The male of I. longipes comb. nov. can be separated from the male of I. rossi by the following combination of features: broad retrolateral tibial apophysis (narrow in I. rossi ), fringed flange with a prolateral twist (fringed flange with retrolateral twist in I. rossi ) and absence of a median apophysis ( I. rossi with highly reduced median apophysis) (compare Figs 10F and 11 A–C with Platnick & Gajbe 1994 : figs 1–3). Females can be separated from the females of I. rossi by an anteriorly oriented long, median tongue-like plate ( Fig. 10F ), which is absent in the latter species ( Gravely 1931 ; Platnick 1991 : fig. 21). FIGURE 9. Sphingius sp.. A–D Ƌ left pedipalp: A prolateral, B ventral; C retrolateral; D enlarged view of retrolateral tibial apophysis, retrolateral. E ♀ epigynum, ventral. Abbreviations: C conductor, E embolus, MA median tegular apophysis, RTA retrolateral tibial apophysis, TA accessory tegular apophysis. Scale bars: A–E, 0.2 mm. Note. With the transfer of S. longipes , the total number of known Inthaeron species is increased to two, I. rossi and I. longipes comb. nov. ; both species are known from both sexes. Remarks. The ZSI collection has one glass bottle for this species labelled as “ Type ” (1513/18), containing a male specimen in good condition, with intact pedipalps. Gravely (1931) also described the female of this species ( Fig. 10F ), but we could not trace any female specimen in the collection.