Revision of the flightless click-beetle genus Dima Charpentier, 1825 (Coleoptera: Elateridae: Dimini) in the Balkan Peninsula
Author
Mertlik, Josef
Pohřebačka 34, CZ- 53345 Opatovice nad Labem, Czech Republic
mertlik@elateridae.com
Author
Németh, Tamás
Hungarian Natural History Museum, Department of Zoology, Baross u. 13, H- 1088 Budapest, Hungary
haesito@gmail.com
Author
Kundrata, Robin
Department of Zoology, Faculty of Science, Palacky University, 17. listopadu 50, CZ- 77146 Olomouc, Czech Republic
robin.kundrata@upol.cz
text
Zootaxa
2017
2017-01-13
4220
1
1
63
journal article
7453
10.11646/zootaxa.4220.1.1
eb597d59-1f6e-412d-a99a-de1a67e7cc94
1175-5326
4670787
D74BC90C-84CC-4788-9048-54F5C8521B32
Dima evritaniensis
Schimmel & Platia, 2008
Figs 26–28
,
126–127
,
183
,
245
.
Dima evritaniensis
Schimmel & Platia, 2008
: 573
.
Type depositories.
Holotype, ♂ (MRSN), 3 paratypes: ♀♀ (MCSB, PCGP, PCRS).
Type
locality.
Greece
:
Evritania province
,
Panetoliko Mts.
,
620 m
, Proussos env.
Redetermined material.
One of the
paratypes
of
D
.
evritaniensis
, a male from Mt. Ossa (
PCGP
;
Fig. 87
), belongs under
D
.
pecoudi
Fleutiaux, 1944
.
New
material.
GREECE
: distr.
Evritania
,
Timfristos Mts.
[Veluchi],
1 ♀
, without further data (
PCVD
)
;
distr.
Evritania
,
Timfristos Mts.
,
Karpeníssi
env., firry wood,
1446 m
(
38°55'25.22"N
,
21°48'33.22"E
), 13.
VI.2012, 4
♂♂,
14 ♀♀
,
J. Mertlik
leg. (
PCJM
)
; dtto, 1 ♂ (PCRK); dtto, 8 ♂♂, 28 ♀♀, V. Dušánek leg. (PCVD); dtto, 3 ♂♂, 11 ♀♀, P. Brůha leg. (PCPB); dtto, 4 ♂♂, 8 ♀♀, B. Zbuzek leg. (PCBZ);
distr.
Evritania
,
Timfristos Mts.
,
Karpeníssi
env., firry wood,
1446 m
(
38°55'25.22"N
,
21°48'33.22"E
), 8.
VI.2015, 8
♂♂,
9 ♀♀
,
J. Mertlik
leg. (
PCJM
)
; dtto, 4 ♂♂, 7 ♀♀, B. Zbuzek leg. (PCBZ); dtto, 2 ♂♂, 14 ♀♀, P. Brůha leg. (PCPB); dtto, 6 ♂♂, 1 ♀, T. Németh leg. (PCRK); dtto, 1 ♂ (HNHM); dtto, 9.
VI.2015, 1
♀, J. Mertlik leg (PCJM).
Diagnosis.
Dima evritaniensis
is a medium-sized species (
11.3–13.5 mm
) with dense pronotal punctuation, semi-erect setae on the pronotal lateral margin, and elytra with short, dense, decumbent pubescence (
Figs 26–28
,
126–127
). This species is morphologically very like
D
.
fthiotidensis
, based on the body shape, size, and matt pronotum with dense, fine punctation (
Figs 35–41
).
Dima fthiotidensis
has slightly coarser punctation of pronotum (
Figs 130–134
), shorter hairs on pronotum and elytra, decumbent to semi-erect pubescence on the pronotal sides (in most cases), and shorter apical lobe of paramera (
Figs 186–187
).
Dima evritaniensis
differs from the remaining species in the
central
Greece
by its matt body surface and pronotum with denser, finer punctation (
Figs 26–28
,
126–127
). Additionally,
D
.
pelionensis
sp. nov.
has the sides of pronotum with decumbent setae in the anterior two thirds (semi-erect setae along its whole length in
D
.
evritaniensis
;
Figs 126–127
,
152–153
),
D
.
etoliensis
has the scutellar frontal margin flattened, declined gradually to the plane of elytra (convex, steeply declined in
D
.
evritaniensis
; visible from the lateral view),
D
.
hladilorum
has more robust apical lobe of paramera (
Fig. 190
), broader scutellum, which is less convex basally (in most cases; scutellum longer and more convex basally in
D
.
evritaniensis
), and
D
.
zbuzeki
sp. nov.
is a smaller species with much longer apical parameral lobe (
Fig. 221
).
Intraspecific variability.
In some cases, elytra or whole body is paler than in a typical form.
Distribution.
Greece
(Evritania: Panetoliko Mts., Timfristos Mts.;
Fig. 245
).
Remarks.
We did not have an opportunity to study the
holotype
of
D
.
evritaniensis
, which is deposited in the Natural History Museum in Torino (MRSN). The collection is closed and not available for study (M. Garsena, pers. comm.). We studied the female
paratype
with the same locality data as the
holotype
(i.e. Panetoliko Mts., Proussos env.; from MCSB;
Fig. 26
) and this is conspecific with the specimens from Timfristos Mts. (
Figs 27–28
). Based on the comparison of the high-resolution
holotype
male photograph (kindly provided us by R. Schimmel) and fresh male specimens collected in Timfristos Mts. in
2012 and 2015
, these populations share similar body size and shape, pronotal punctation and pubescence, and length of antennae. According to the original description of
D
.
evritaniensis
(
Schimmel & Platia 2008
)
, the
holotype
has short antennomeres II and III, with antennomere III slightly longer than II, and both combined as long as antennomere IV. However, all specimens which we examined here (including the
paratype
) have antennomere II slightly elongate, subtriangular, narrower than antennomere III, antennomeres II and III of subequal length, both combined longer than antennomere IV. Additionally, the parameral apical lobes of
D. evritaniensis
figured in the original description (Schimmel & Platia 2 008) are shorter than in the specimens from Timfristos, however, it might be caused by the different angle and/or the low-quality photograph. Therefore, the real shape of paramera might differ from the one originally figured (compare e.g. the shape of paramera of
D
.
assingi
holotype
in the original description and herein;
Schimmel & Platia (2008)
;
Fig. 175
).