Evolution of the hectocotylus in Sepiolinae (Cephalopoda: Sepiolidae) and description of four new genera Author Bello, Giambattista text European Journal of Taxonomy 2020 2020-05-29 655 1 53 journal article 21713 10.5852/ejt.2020.655 fadee126-7da7-47d9-b9ac-973697ecf9f5 2118-9773 3888797 0042EFAE-2E4F-444B-AFB9-E321D16116E8 Subfamily Sepiolinae Leach, 1817 Sepiolinae Leach, 1817: 137 ; nom. transl. ex Appellöf, 1898. Type genus: Sepiola Leach, 1817 . Diagnosis Mantle broad, U-shaped, dorsally fused with head in nuchal region by occipital band; without ventral mantle shield. Fins rounded or slightly pointed; their length to about half mantle length or slightly greater, posteriorly not reaching the mantle rear end. Interbrachial web weakly developed between arms I–III. Arm suckers in 2 or 4, exceptionally 8, longitudinal rows. Left dorsal arm of male hectocotylized. Bursa copulatrix present near female genital opening in the left ventral side of mantle cavity. Visceral photophores present or absent. Gladius reduced or absent. (Modified after Vecchione & Young 2004 and Reid & Jereb 2005).