Ultrastructure of attachment specializations of hexapods (Arthropoda): evolutionary patterns inferred from a revised ordinal phylogeny Author R. G. Beutel Author S. N. Gorb text J. Zool. Syst. Evol. Research 2001 39 177 207 journal article 10.1046/j.1439-0469.2001.00155.x 95b3d62f-8fcc-4088-873f-b33f4a63d3fc 2360323 3 Diplura ( Grassé 1949 ; Condé and Pagés 1991 ) Presence of an empodial median claw and of a pair of larger, lateral claws is probably a groundplan feature of the order. This condition is found in Anajapyx and Lepidocampa species. The median claw is lost in all other diplurans.