The Southeast Asian Pholcus halabala species group (Araneae, Pholcidae): new data from field observations and ultrastructure
Author
Berhard A. Huber
Author
Booppa Petchard
Author
Charles Leh Moi Ung
Author
Joseph K. H. Koh
Author
Amir R. M. Ghazali
text
European Journal of Taxonomy
2016
190
1
55
http://www.europeanjournaloftaxonomy.eu/index.php/ejt/article/view/313
journal article
32674
10.5852/ejt.2016.190
3c438ae3-9ea0-4eda-b893-acf6d5756c90
831004
urn:lsid:zoobank.org:pub:BE92596B-62D9-46CD-8486-CF6B36C640B11
Pholcus sudhami
Huber, 2011
Figs 54–56
,
79–89
Pholcus sudhami
Huber, 2011
: 300
, figs
1380–1382
,
1411–1412
,
1476–1480
(
♂♀
).
Diagnosis
Easily distinguished from most congeners by male palpal morphology (figs 1476, 1477 in
Huber 2011
; distinctive procursus with large membranous structure, absence of appendix) and by female genitalia (figs
1479 and
1480
in
Huber 2011
; elongate pore plates in lateral position and semicircular internal structures visible through cuticle; very similar to
Ph. pakse
); from
Ph. pakse
only by male embolus without spines (compare figs
1476 and
1481
in
Huber 2011
).
Figs 79–89.
Pholcus sudhami
Huber, 2011
, ZFMK Ar 15031–32.
79–80
. Male and female prosomata, frontal views.
81
. Female ALS.
82
. Comb-hairs on male tarsus 4.
83
. Right procursus, retrolateral (slightly distal) view.
84
. Left bulb and procursus, prolateral view.
85
. Left uncus and embolus, prolateral view.
86
. Male distal cheliceral apophysis (arrows point at modified hairs).
87
. Epigynum, ventral view.
88
. Male gonopore.
89
. Pseudotrichia on ventral side of procursus. Abbreviations: b = genital bulb; e = embolus; p = procursus; u = uncus. Scale lines: 20 µm (81–82, 86); 30 µm (89); 50 µm (88); 60 µm (85); 200 µm (83–84, 87); 300 µm (79–80).
Figs 90–101.
Female genitalia, untreated in ventral view, cleared in ventral and dorsal views.
90– 92
.
Pholcus
ubin
Huber
,
sp. nov.
93–95
.
Ph.
lintang
Huber
,
sp. nov.
96–98
.
Ph.
kuhapimuk
Huber
,
sp. nov.
99–101
.
Ph.
khaolek
Huber
,
sp. nov.
New material examined
THAILAND
:
9 ♂♂
,
8 ♀♀
,
ZFMK
(
8 ♂♂
,
7 ♀♀
,
Ar 15031–32
)
and
PSUZC
(
1 ♂
,
1 ♀
),
Kanchanaburi
,
Erawan
National Park
(
14°22.2' N
,
99°08.75' E
),
85 m
a.s.l.,
forest along stream, on rocks and tree roots
,
15 Mar. 2015
(
B.A. Huber
,
B. Petcharad
)
;
2 ♀♀
, in absolute ethanol,
ZFMK
(
Mal 379
), same data
;
1 ♂
,
2 ♀♀
,
RMNH
,
Erawan
National Park
,
15–16 Mar. 1986
(
C.L. & P.R. Deeleman
)
.
Assigned tentatively
THAILAND
:
3 ♂♂
,
3 ♀♀
,
ZFMK
(
Ar 15033
),
Prachuap Khiri Khan
,
Khao Sam Roi Yot National Park
,
near Tham Phraya Nakhon
(
12°12.0' N
,
100°00.8' E
),
40 m
a.s.l.,
at rocks in forest
,
14 Mar. 2015
(
B.A. Huber
,
B. Petcharad
)
;
2 ♀♀
,
3 juvs
, in absolute ethanol,
ZFMK
(
Mal 373
), same data
;
1 ♀
,
ZFMK
(
Ar 15034
),
Khao Sam Roi Yot National Park
,
Tham Kaeo
(
12°12.2' N
,
99°59.5' E
),
60 m
a.s.l.,
under rock in cave entrance
,
14 Mar. 2015
(
B.A. Huber
,
B. Petcharad
)
;
1 ♀
,
RMNH
,
Sam Roi Yot National Park
, no further locality information,
8 Dec. 1990
(
C.L. & P.R. Deeleman
)
;
1 juv.
, in absolute ethanol,
ZFMK
(
Mal 376
),
Wat Huai Takaeng
(
13°35.23' N
,
99°45.52' E
) (= “
Tham Phraya Prap
” in
Huber
2011
),
30 m
a.s.l.,
in cave
,
15 Mar. 2015
(
B.A. Huber
,
B. Petcharad
)
.
Description – amendments
In males from Khao Sam Roi Yot, the uncus has a slightly more pointed tip and the subdistal spine on the embolus has a more transversal position relative to the embolus. Specimens from this locality are therefore assigned tentatively. Tibia
1 in
7 males
from
type
locality (
Erawan
): 8.7–10.6 (mean 9.9); in
7 females
: 7.5–8.6 (mean 8.1). Hairs on male ocular area not visibly different from those in females (
Figs 79–80
); tarsus 4 comb-hairs of the simplified
Pholcus
-
type
(cf.
Huber & Fleckenstein 2008
), with three lateral tines (
Fig. 82
); procursus with distinctive field of ventral spines (
Figs 83, 89
); male distal cheliceral apophyses with two modified hairs each (
Fig. 86
); gonopore with four epiandrous spigots (
Fig. 88
); ALS with one widened, one pointed, and six smaller cylindrically shaped spigots of varying sizes (
Fig. 81
).
Natural history
At the
type
locality (
Erawan
) most specimens were found on the lower surfaces of exposed tree roots at the riverside. At the other localities, specimens were only found on overhanging smooth rocks at the cave entrances (Tham Kaeo; Wat Huai Takaeng) or in the forest (Phraya Nakhon). When disturbed, the spiders dropped to the ground.
Distribution
Known from four localities in central western Thailand (
Fig. 57
).