New records and description of four new species of spionids (Annelida: Polychaeta: Spionidae) from the Philippines: the genera Dispio, Malacoceros, Polydora, and Scolelepis, with notes on palp ciliation patterns of the genus Scolelepis Author Williams, Jason D. text Zootaxa 2007 1459 1 35 journal article 10.5281/zenodo.176375 4ae7ea7a-e406-4c68-9777-ec29e3521997 1175-5326 176375 Key to the Spionidae of the Philippines * 1. Specialized setae present on setiger 5..........................................................................................................7 - Specialized setae absent on setiger 5 ........................................................................................................... 2 2. Prostomium pointed..................................................................................................................................... 3 - Prostomium with lateral horns ............................................................. Malacoceros indicus ( Fauvel, 1928 ) 3. Branchiae from setiger 1; with accessory branchiae ............................................. Dispio latilamella sp. n. - Branchiae from setiger 2; without accessory branchiae .............................................................................. 4 4. Occipital tentacle present .............................................................................. Scolelepis magnicornuta sp. n. - Occipital tentacle absent.............................................................................................................................. 5 5. Notopodial lamellae of anterior setigers with up to nine conical lobes................. Scolelepis alisonae sp. n. - Notopodial lamellae of anterior setigers without conical lobes................................................................... 6 6. Notopodial hooded hooks begin on same setigers as neuropodial hooded hooks......................................... .............................................................................................................. Scolelepis hutchingsae Dauer, 1985 - Notopodial hooded hooks begin at least ten setigers posterior to beginning of neuropodial hooded hooks. ......................................................................................................................... Scolelepis villosivaina sp. n. 7. Setiger 5 with one type of major spine ........................................................................................................ 8 - Setiger 5 with two types of major spines................................................................................................... 16 8. Branchiae from setiger 2; tridentate hooded hooks present; setiger 4 with modified notosetae; setiger 5 only weakly modified ........................................................................ Tripolydora spinosa Woodwick, 1964 - Branchiae begin posterior to setiger 5, or absent; bidentate hooded hooks; setiger 5 highly modified ...... 9 9. Notosetae present on setiger 1; hooded hooks without constriction on shaft ..................... Dipolydora ….10 - Notosetae absent on setiger 1; hooded hooks with constriction on shaft ............................. Polydora ….12 10. Major spines of setiger 5 bidentate with a large lateral tooth and cowling connecting teeth; posterior noto- podia with awl-like modified spines ............................................... Dipolydora armata ( Langerhans, 1880 ) - Posterior notopodia lacking modified spines............................................................................................ 11 11. Major spines of setiger 5 with small subterminal protuberance on convex side; pygidium with large ven- tral lobe and 2 smaller dorsal lobes .................................................. Dipolydora socialis ( Schmarda, 1861 ) - Major spines of setiger 5 with two lateral teeth; small cuff-shaped pygidium.............................................. ............................................................................................... Dipolydora tridenticulata ( Woodwick, 1964 ) 12. Bundles of needle-like notosetae protruding through cuticle of posterior setigers; pygidium with anal cirri ....................................................................................................................... Polydora robi Williams, 2000 - Bundles of posterior notosetae absent; pygidium cup-shaped................................................................... 13 13. Occipital tentacle absent............................................................................................................................ 14 - Occipital tentacle present ............................................................................ Polydora cavitensis Pillai, 1965 14. Palps without pigmentation ....................................................................................................................... 15 - Palps with bands of black pigment ......................................................... Polydora sp. A Williams, 2001** 15. Major spines of setiger 5 simple, falcate .................................................. Polydora mabinii Williams, 2001 - Major spines of setiger 5 with large lateral tooth .............................. Polydora umangivora Williams, 2001 16. Branchiae from setiger 2.............................................. Boccardia berkeleyorum Blake & Woodwick, 1971 - Branchiae from setiger 7, or absent ........................................................................................................... 17 17. Up to ten pairs of branchiae beginning on setiger 7; a borer in calcareous substrata.................................... ...................................................................................................... Carazziella reishi ( Woodwick, 1964 ) *** - Branchiae reduced, one or two pairs beginning on setiger 7 or absent; associated with sponges............ 18 18. Unilimbate capillary neurosetae in setigers 2–7; setiger 5 modified with two types of major spines arranged in a double row: dorsal row falcate with an apical shelf; ventral row with expanded distal end bearing up to 9 teeth along the apical edge and a capillary extension on the posterior end.......................... .................................................................................................... Polydorella dawydoffi Radashevsky, 1996 - Acicular neurosetae in setigers 2–7; setiger 5 modified with two types of major spines arranged in a dou- ble row: dorsal row simple, falcate; ventral row with slightly expanded distal end bearing small digitiform bosses .............................................................................................. Polydorella kamakamai Williams, 2004