<document id="8DD744959F1C02DDA86192A2FFA593B5" ID-DOI="10.5281/zenodo.11404264" ID-ISSN="1525-9153" ID-Zenodo-Dep="11404264" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="juliana" IM.tables_approvedBy="juliana" IM.taxonomicNames_approvedBy="juliana" IM.treatments_approvedBy="juliana" checkinTime="1716907766958" checkinUser="felipe" docAuthor="Barrio-Amorós, César L., Rojas-Runjaic, Fernando J. M. &amp; Señaris, J. Celsa" docDate="2019" docId="4608879FFFD4FFE16E1BFCB2FDB2F8EE" docLanguage="en" docName="AmphibReptileCons.13.1.e180.pdf" docOrigin="Amphibian &amp; Reptile Conservation (e 180) 13 (1)" docStyle="DocumentStyle:2DCF38F802BD4DFFCCCB5916C58E77B3.2:AmphibReptileConse.2015-.journal_article.2cover" docStyleId="2DCF38F802BD4DFFCCCB5916C58E77B3" docStyleName="AmphibReptileConse.2015-.journal_article.2cover" docStyleVersion="2" docTitle="Pristimantis memorans" docType="treatment" docVersion="5" lastPageNumber="120" masterDocId="BA31FFE7FFACFF986D36FA1CFFB9FFDF" masterDocTitle="Catalogue of the amphibians of Venezuela: Illustrated and annotated species list, distribution, and conservation" masterLastPageNumber="198" masterPageNumber="1" pageNumber="119" updateTime="1717186961153" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="C691AEB0FD42EA8942923BEEC0EDF071">Catalogue of the amphibians of Venezuela: Illustrated and annotated species list, distribution, and conservation</mods:title>
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<mods:namePart id="B2C5AB2F1FBC04545EA9CBDCDC04A9A9">Barrio-Amorós, César L.</mods:namePart>
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<mods:namePart id="CCC04E3AFB52B2E20A6AE24B24B04C8D">Rojas-Runjaic, Fernando J. M.</mods:namePart>
<mods:affiliation id="1C7C595063B88638DD2160142FD01407">Fundación La Salle de Ciencias Naturales, Museo de Historia Natural La Salle, Apartado Postal 1930, Caracas 1010 - A, VENEZUELA &amp; Current address: Pontifícia Universidade Católica do Río Grande do Sul (PUCRS), Laboratório de Sistemática de Vertebrados, Av. Ipiranga 6681, Porto Alegre, RS 90619 - 900, BRAZIL</mods:affiliation>
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<mods:namePart id="40FAC1C666570C6F4A3F65E3C1D50773">Señaris, J. Celsa</mods:namePart>
<mods:affiliation id="F7037BA9383021CDB7856D3081B83019">Instituto Venezolano de Investigaciones Científicas, Altos de Pipe, apartado 20632, Caracas 1020, VENEZUELA</mods:affiliation>
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<mods:title id="02C2170593437DE603E6D70BB35125BF">Amphibian &amp; Reptile Conservation</mods:title>
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<treatment id="4608879FFFD4FFE16E1BFCB2FDB2F8EE" ID-DOI="http://doi.org/10.5281/zenodo.11405926" ID-Zenodo-Dep="11405926" LSID="urn:lsid:plazi:treatment:4608879FFFD4FFE16E1BFCB2FDB2F8EE" httpUri="http://treatment.plazi.org/id/4608879FFFD4FFE16E1BFCB2FDB2F8EE" lastPageId="121" lastPageNumber="120" pageId="120" pageNumber="119">
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<heading id="955681E5FFD4FFE06E1BFCB2FAE4F919" box="[813,1373,1710,1734]" centered="true" fontSize="10" level="2" pageId="120" pageNumber="119" reason="4">
<taxonomicName id="09A14D0AFFD4FFE06E1BFCB2FAE4F919" ID-CoL="77WG5" authority="(Myers and Donnelly 1997)" baseAuthorityName="Myers and Donnelly" baseAuthorityYear="1997" box="[813,1373,1710,1734]" class="Amphibia" family="Craugastoridae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="120" pageNumber="119" phylum="Chordata" rank="species" species="memorans">
<emphasis id="FCD5EA9BFFD4FFE06E1BFCB2FB94F91A" bold="true" box="[813,1069,1710,1734]" italics="true" pageId="120" pageNumber="119">Pristimantis memorans</emphasis>
(
<bibRefCitation id="AA304B78FFD4FFE0690DFCB2FAECF919" author="Myers CW &amp; Donnelly MA" box="[1083,1365,1710,1734]" pageId="120" pageNumber="119" pagination="1 - 71" refId="ref102658" refString="Myers CW, Donnelly MA. 1997. A tepui herpetofauna on a granitic mountain (Tamacuari) in the borderland between Venezuela and Brazil. Report of the Phipps- Tapirapeco expedition. American Museum Novitates 3213: 1 - 71." type="journal article" year="1997">Myers and Donnelly 1997</bibRefCitation>
)
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
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<materialsCitation id="7EC93CD4FFD4FFE06E21FCF2FB85F8D9" box="[791,1084,1774,1798]" collectionCode="AMNH" pageId="120" pageNumber="119" specimenCode="AMNH 131466" specimenCount="1" typeStatus="holotype">
<emphasis id="FCD5EA9BFFD4FFE06E21FCF2FC3EF8D9" bold="true" box="[791,903,1774,1798]" pageId="120" pageNumber="119">
<typeStatus id="111A882BFFD4FFE06E21FCF2FC3AF8D9" box="[791,899,1774,1798]" pageId="120" pageNumber="119" type="holotype">Holotype</typeStatus>
:
</emphasis>
<specimenCode id="9E079EF2FFD4FFE06EBAFCF2FB81F8D9" box="[908,1080,1774,1798]" collectionCode="AMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="120" pageNumber="119">AMNH 131466</specimenCode>
.
</materialsCitation>
</paragraph>
<paragraph id="CE1E3689FFD4FFE06E21FD12FC40F8B9" blockId="120.[791,1395,1774,1958]" pageId="120" pageNumber="119">
<emphasis id="FCD5EA9BFFD4FFE06E21FD12FC02F8FA" bold="true" box="[791,955,1805,1830]" pageId="120" pageNumber="119">Type locality:</emphasis>
“ 
<materialsCitation id="7EC93CD4FFD4FFE06EE3FD12FC40F8B9" country="Venezuela" elevation="1" latitude="1.2166667" location="Sierra Tapirapeco" longLatPrecision="1308" longitude="-64.7" pageId="120" pageNumber="119" specimenCount="1" stateProvince="Amazonas">
Ridge N Pico Tamacuari, 
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<elevation id="458CD1BAFFD4FFE06827FD12FACAF8FA" box="[1297,1395,1806,1830]" metricMagnitude="0" metricUnit="m" metricValue="1.27" pageId="120" pageNumber="119" unit="m" value="1.27">1,270 m</elevation>
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elevation, 
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, 
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, 
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(
<geoCoordinate id="AB95504EFFD4FFE06E16FD52FCC9F8B9" box="[800,880,1870,1894]" degrees="1" direction="north" minutes="13" orientation="latitude" pageId="120" pageNumber="119" precision="925" value="1.2166667">1°13’N</geoCoordinate>
, 
<geoCoordinate id="AB95504EFFD4FFE06E4AFD52FC5BF8B9" box="[892,994,1870,1894]" degrees="64" direction="west" minutes="42" orientation="longitude" pageId="120" pageNumber="119" precision="925" value="-64.7">64°42’W</geoCoordinate>
).”
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</paragraph>
</subSubSection>
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<paragraph id="CE1E3689FFD4FFE16E21FD71FD6DF9AE" blockId="120.[791,1395,1774,1958]" lastBlockId="121.[141,746,1625,1842]" lastPageId="121" lastPageNumber="120" pageId="120" pageNumber="119">
<emphasis id="FCD5EA9BFFD4FFE06E21FD71FC12F85A" bold="true" box="[791,939,1901,1925]" pageId="120" pageNumber="119">Distribution:</emphasis>
Region 5. Known from 
<typeStatus id="111A882BFFD4FFE0698AFD72FB53F859" box="[1212,1258,1902,1926]" pageId="120" pageNumber="119">type</typeStatus>
locality and one additional locality, a distance of 
<quantity id="09599B6CFFD4FFE06995FD91FB51F87A" box="[1187,1256,1933,1957]" metricMagnitude="2" metricUnit="m" metricValue="3.5" pageId="120" pageNumber="119" unit="m" value="350.0">350 m</quantity>
on Brazilian side of Tapirapecó (
<bibRefCitation id="AA304B78FFD5FFE16C69FC46FD71F9AE" author="Caramaschi U &amp; Niemeyer H." box="[351,712,1625,1650]" pageId="121" pageNumber="120" pagination="199" refId="ref88106" refString="Caramaschi U, Niemeyer H. 2005 c. Geographic distribution: Eleutherodactylus cavernibardus. Herpetological Review 36: 199." type="journal article" year="2005">Caramaschi and Niemeyer 2005c</bibRefCitation>
).
</paragraph>
</subSubSection>
<caption id="9ADE6601FFD5FFE16DB8F855FE05FDA3" ID-DOI="http://doi.org/10.5281/zenodo.11405023" ID-Zenodo-Dep="11405023" httpUri="https://zenodo.org/record/11405023/files/figure.png" pageId="121" pageNumber="120" startId="121.[142,178,585,606]" targetBox="[166,1395,141,576]" targetPageId="121" targetType="figure">
<paragraph id="CE1E3689FFD5FFE16DB8F855FE05FDA3" blockId="121.[142,747,585,636]" pageId="121" pageNumber="120">
<emphasis id="FCD5EA9BFFD5FFE16DB8F855FF71FD81" bold="true" box="[142,200,585,606]" pageId="121" pageNumber="120">256A.</emphasis>
<emphasis id="FCD5EA9BFFD5FFE16DEEF855FE11FD81" box="[216,424,585,606]" italics="true" pageId="121" pageNumber="120">
<taxonomicName id="09A14D0AFFD5FFE16DEEF855FE1DFD81" baseAuthorityName="Lutz" baseAuthorityYear="1927" box="[216,420,585,606]" class="Amphibia" family="Craugastoridae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="121" pageNumber="120" phylum="Chordata" rank="species" species="incertus">Pristimantis incertus</taxonomicName>
.
</emphasis>
Holotype of 
<taxonomicName id="09A14D0AFFD5FFE16F76F855FF65FDA3" authority=", Original" authorityName="Original" class="Amphibia" family="Hylodidae" genus="Hylodes" kingdom="Animalia" order="Anura" pageId="121" pageNumber="120" phylum="Chordata" rank="species" species="incertus">
<emphasis id="FCD5EA9BFFD5FFE16F76F855FD5DFD81" box="[576,740,585,606]" italics="true" pageId="121" pageNumber="120">Hylodes incertus</emphasis>
, Original
</taxonomicName>
drawing in 
<bibRefCitation id="AA304B78FFD5FFE16C7EF87AFE01FDA3" author="Lutz A." box="[328,440,614,636]" pageId="121" pageNumber="120" pagination="35 - 65" refId="ref99402" refString="Lutz A. 1927. Notas sobre batrachios da Venezuela e da Ilha de trinidad. Memorias do Instituto Oswaldo Cruz 20: 35 - 65." type="journal article" year="1927">Lutz (1927)</bibRefCitation>
.
</paragraph>
</caption>
<caption id="9ADE6601FFD5FFE16E21F855FBE0FDA4" pageId="121" pageNumber="120" startId="121.[791,828,585,606]" targetType="figure">
<paragraph id="CE1E3689FFD5FFE16E21F855FBE0FDA4" blockId="121.[791,1395,585,635]" pageId="121" pageNumber="120">
<emphasis id="FCD5EA9BFFD5FFE16E21F855FCF6FD81" bold="true" box="[791,847,585,606]" pageId="121" pageNumber="120">256B.</emphasis>
<emphasis id="FCD5EA9BFFD5FFE16E6DF855FB90FD81" box="[859,1065,585,606]" italics="true" pageId="121" pageNumber="120">
<taxonomicName id="09A14D0AFFD5FFE16E6DF855FB9CFD81" baseAuthorityName="Lutz" baseAuthorityYear="1927" box="[859,1061,585,606]" class="Amphibia" family="Craugastoridae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="121" pageNumber="120" phylum="Chordata" rank="species" species="incertus">Pristimantis incertus</taxonomicName>
.
</emphasis>
Dorsal view of the holotype of 
<taxonomicName id="09A14D0AFFD5FFE16E21F87AFBECFDA4" authority=", AL-MNRJ 1190" authorityName=", AL-MNRJ" authorityYear="1190" box="[791,1109,614,635]" class="Amphibia" family="Hylodidae" genus="Hylodes" kingdom="Animalia" order="Anura" pageId="121" pageNumber="120" phylum="Chordata" rank="species" species="incertus">
<emphasis id="FCD5EA9BFFD5FFE16E21F87AFC09FDA4" box="[791,944,614,635]" italics="true" pageId="121" pageNumber="120">Hylodes incertus</emphasis>
, AL-MNRJ 1190
</taxonomicName>
.
</paragraph>
</caption>
<caption id="9ADE6601FFD5FFE16DBBFE69FEC6FB77" ID-DOI="http://doi.org/10.5281/zenodo.11405025" ID-Zenodo-Dep="11405025" httpUri="https://zenodo.org/record/11405025/files/figure.png" pageId="121" pageNumber="120" startId="121.[141,178,1141,1163]" targetBox="[141,1395,648,1131]" targetPageId="121" targetType="figure">
<paragraph id="CE1E3689FFD5FFE16DBBFE69FEC6FB77" blockId="121.[141,746,1141,1192]" pageId="121" pageNumber="120">
<emphasis id="FCD5EA9BFFD5FFE16DBBFE69FF7FFB54" bold="true" box="[141,198,1141,1163]" pageId="121" pageNumber="120">256C.</emphasis>
<emphasis id="FCD5EA9BFFD5FFE16DFAFE6AFE2BFB54" box="[204,402,1142,1163]" italics="true" pageId="121" pageNumber="120">
<taxonomicName id="09A14D0AFFD5FFE16DFAFE6AFE37FB54" baseAuthorityName="Lutz" baseAuthorityYear="1927" box="[204,398,1142,1163]" class="Amphibia" family="Craugastoridae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="121" pageNumber="120" phylum="Chordata" rank="species" species="incertus">Pristimantis incertus</taxonomicName>
.
</emphasis>
Left hand of the holotype of 
<taxonomicName id="09A14D0AFFD5FFE16FA8FE6AFEC2FB77" authority=", AL-MNRJ 1190" authorityName=", AL-MNRJ" authorityYear="1190" class="Amphibia" family="Hylodidae" genus="Hylodes" kingdom="Animalia" order="Anura" pageId="121" pageNumber="120" phylum="Chordata" rank="species" species="incertus">
<emphasis id="FCD5EA9BFFD5FFE16FA8FE6AFF6FFB77" italics="true" pageId="121" pageNumber="120">Hylodes incertus</emphasis>
, AL-MNRJ 1190
</taxonomicName>
.
</paragraph>
</caption>
<caption id="9ADE6601FFD5FFE16E21FE69FB3DFB77" pageId="121" pageNumber="120" startId="121.[791,827,1141,1162]" targetType="figure">
<paragraph id="CE1E3689FFD5FFE16E21FE69FB3DFB77" blockId="121.[791,1396,1141,1192]" pageId="121" pageNumber="120">
<emphasis id="FCD5EA9BFFD5FFE16E21FE69FCE9FB55" bold="true" box="[791,848,1141,1162]" pageId="121" pageNumber="120">256D.</emphasis>
<emphasis id="FCD5EA9BFFD5FFE16E61FE6AFBA7FB54" box="[855,1054,1142,1163]" italics="true" pageId="121" pageNumber="120">
<taxonomicName id="09A14D0AFFD5FFE16E61FE6AFBA2FB54" baseAuthorityName="Lutz" baseAuthorityYear="1927" box="[855,1051,1142,1163]" class="Amphibia" family="Craugastoridae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="121" pageNumber="120" phylum="Chordata" rank="species" species="incertus">Pristimantis incertus</taxonomicName>
.
</emphasis>
Male. Plain phase, Cerro El Volcán, Miranda. Photo: César Barrio-Amorós.
</paragraph>
</caption>
<caption id="9ADE6601FFD5FFE16DBBFC1AFE76F9E7" ID-DOI="http://doi.org/10.5281/zenodo.11405029" ID-Zenodo-Dep="11405029" httpUri="https://zenodo.org/record/11405029/files/figure.png" pageId="121" pageNumber="120" startId="121.[141,177,1542,1563]" targetBox="[141,1394,1204,1535]" targetPageId="121" targetType="figure">
<paragraph id="CE1E3689FFD5FFE16DBBFC1AFE76F9E7" blockId="121.[141,747,1542,1593]" pageId="121" pageNumber="120">
<emphasis id="FCD5EA9BFFD5FFE16DBBFC1AFF7CF9C4" bold="true" box="[141,197,1542,1563]" pageId="121" pageNumber="120">256E.</emphasis>
<emphasis id="FCD5EA9BFFD5FFE16DE7FC1BFE27F9C3" box="[209,414,1543,1564]" italics="true" pageId="121" pageNumber="120">
<taxonomicName id="09A14D0AFFD5FFE16DE7FC1BFE23F9C3" baseAuthorityName="Lutz" baseAuthorityYear="1927" box="[209,410,1543,1564]" class="Amphibia" family="Craugastoridae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="121" pageNumber="120" phylum="Chordata" rank="species" species="incertus">Pristimantis incertus</taxonomicName>
.
</emphasis>
Patterned phase. Cerro Zapatero, Yaracuy. Photo: Sebastian Lotzkat.
</paragraph>
</caption>
<caption id="9ADE6601FFD5FFE16E21FC1AFBE0F9E7" pageId="121" pageNumber="120" startId="121.[791,826,1542,1563]" targetType="figure">
<paragraph id="CE1E3689FFD5FFE16E21FC1AFBE0F9E7" blockId="121.[791,1396,1542,1593]" pageId="121" pageNumber="120">
<emphasis id="FCD5EA9BFFD5FFE16E21FC1AFCF2F9C4" bold="true" box="[791,843,1542,1563]" pageId="121" pageNumber="120">256F.</emphasis>
<emphasis id="FCD5EA9BFFD5FFE16E6FFC1BFB91F9C3" box="[857,1064,1543,1564]" italics="true" pageId="121" pageNumber="120">
<taxonomicName id="09A14D0AFFD5FFE16E6FFC1BFB9DF9C3" baseAuthorityName="Lutz" baseAuthorityYear="1927" box="[857,1060,1543,1564]" class="Amphibia" family="Craugastoridae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="121" pageNumber="120" phylum="Chordata" rank="species" species="incertus">Pristimantis incertus</taxonomicName>
.
</emphasis>
Stripped phase. Cerro Zapatero, Yaracuy. Photo: Sebastian Lotzkat.
</paragraph>
</caption>
<subSubSection id="86BB6502FFD5FFE16DBBFC66FDB2F8EE" pageId="121" pageNumber="120" type="discussion">
<paragraph id="CE1E3689FFD5FFE16DBBFC66FDFEF92D" blockId="121.[141,746,1625,1842]" pageId="121" pageNumber="120">
<emphasis id="FCD5EA9BFFD5FFE16DBBFC66FF47F94D" bold="true" box="[141,254,1658,1682]" pageId="121" pageNumber="120">Remarks:</emphasis>
In 
<taxonomicName id="09A14D0AFFD5FFE16C07FC66FDF8F94E" baseAuthorityName="Gunther" baseAuthorityYear="1859" box="[305,577,1658,1681]" class="Amphibia" family="Craugastoridae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="121" pageNumber="120" phylum="Chordata" rank="species" species="unistrigatus">
<emphasis id="FCD5EA9BFFD5FFE16C07FC66FDF8F94E" box="[305,577,1658,1681]" italics="true" pageId="121" pageNumber="120">Pristimantis unistrigatus</emphasis>
</taxonomicName>
species group according to original publication (
<bibRefCitation id="AA304B78FFD5FFE16F30FC86FF7DF90D" author="Myers CW &amp; Donnelly MA" pageId="121" pageNumber="120" pagination="1 - 71" refId="ref102658" refString="Myers CW, Donnelly MA. 1997. A tepui herpetofauna on a granitic mountain (Tamacuari) in the borderland between Venezuela and Brazil. Report of the Phipps- Tapirapeco expedition. American Museum Novitates 3213: 1 - 71." type="journal article" year="1997">Myers and Donnelly 1997</bibRefCitation>
) and by 
<bibRefCitation id="AA304B78FFD5FFE16C1FFCA6FDB2F90D" author="Hedges SB &amp; Duellman WE &amp; Heinicke MP" box="[297,523,1722,1746]" pageId="121" pageNumber="120" pagination="1 - 182" refId="ref94193" refString="Hedges SB, Duellman WE, Heinicke MP. 2008. New World direct-developing frogs (Anura: Terrarana): molecular phylogeny, classification, biogeography, and conservation. Zootaxa 1737: 1 - 182." type="journal article" year="2008">Hedges et al. (2008)</bibRefCitation>
, but not assigned to any species group by 
<bibRefCitation id="AA304B78FFD5FFE16C42FCC6FDFAF92D" author="Padial JM &amp; Grant T &amp; Frost D." box="[372,579,1754,1778]" pageId="121" pageNumber="120" pagination="1 - 132" refId="ref103173" refString="Padial JM, Grant T, Frost D. 2014. Molecular systematics of terraranas (Anura; Brachycephaloidea) with an assessment of the effects of alignment and optimality criteria. Zootaxa 3825: 1 - 132." type="journal article" year="2014">Padial et al. (2014)</bibRefCitation>
.
</paragraph>
<paragraph id="CE1E3689FFD5FFE16DBBFCE5FDB2F8EE" blockId="121.[141,746,1625,1842]" pageId="121" pageNumber="120">
<emphasis id="FCD5EA9BFFD5FFE16DBBFCE5FEC5F8CE" bold="true" box="[141,380,1785,1809]" pageId="121" pageNumber="120">Selected references:</emphasis>
<bibRefCitation id="AA304B78FFD5FFE16CA6FCE6FD5CF8CD" author="Myers CW &amp; Donnelly MA" box="[400,741,1786,1810]" pageId="121" pageNumber="120" pagination="1 - 71" refId="ref102658" refString="Myers CW, Donnelly MA. 1997. A tepui herpetofauna on a granitic mountain (Tamacuari) in the borderland between Venezuela and Brazil. Report of the Phipps- Tapirapeco expedition. American Museum Novitates 3213: 1 - 71." type="journal article" year="1997">Myers and Donnelly (1997)</bibRefCitation>
; 
<bibRefCitation id="AA304B78FFD5FFE16DBBFD06FDBEF8EE" author="Caramaschi U &amp; Niemeyer H." box="[141,519,1817,1842]" pageId="121" pageNumber="120" pagination="74" refId="ref88127" refString="Caramaschi U, Niemeyer H. 2005 d. Geographic distribution: Eleutherodactylus memorans. Herpetological Review 36: 74." type="journal article" year="2005">Caramaschi and Niemeyer (2005d)</bibRefCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>