The millipede family Conotylidae in northwestern North America, with a complete bibliography of the family (Diplopoda, Chordeumatida, Heterochordeumatidea, Conotyloidea)
Author
Shear, William A.
Author
Richart, Casey H.
Author
Wong, Victoria L.
text
Zootaxa
2020
2020-03-19
4753
1
1
78
journal article
21316
10.11646/zootaxa.4753.1.1
f7974bcf-11e7-4d1b-b09d-530dc2dbd54f
1175-5326
3983782
AA9F66B3-EF8C-4F6B-8F35-0BCBEE5122ED
Brunsonia selwayana
,
new species
Figs. 173–179
,
190, 191
Types
:
Male
holotype
and
female
paratype
from
IDAHO
:
Idaho Co
.
,
O’Hara Campground
,
Selway River Road
6.7 mi
SE of
US-12,
Nez Perce-Clearwater National Forests
, elev.
475 m
.
,
46.0845°N
, -
115.5154°W
, collected
12 April 2003
, by
W. Leonard
, from the forest litter of a mixed forest including
Thuja plicata
,
Pseudotsuga menziesii
,
Alnus
cf.
rubra
,
Populus trichocarpa
,
Rubus parviflorus
, and moss, (
CAS
)
.
Male
paratype
, same as previous, collected
14 October 2006
, by W. Leonard, C. Richart, A. Fusek (CAS)
.
Diagnosis
: The posterior gonopod colpocoxites are unique and appear somewhat reduced compared to other
Brunsonia
.
Both this species and the following are distinct from the others in having a prominent, oblong femoral process on male legpair five, while pairs three, six and seven lack femoral processes. The anterior gonopods of
B. benewah
,
n. sp.
have a posterior process that is lacking in the present species.
FIGS. 173–189.
Brunsonia
spp. Figs. 173–179.
Brunsonia chelanomagna
.
Fig. 173. Right anterior gonopod, lateral view. Fig. 174. Left posterior gonopod coxite, posterior view. Figs. 175–179. Male legs 3–7, respectively. Figs. 180, 181.
Brunsonia selwayana
.
Fig. 180. Right anterior gonopod, posteriolateral view. Fig. 181. Left posterior gonopod coxite, posterior view.
Figs. 182–189.
Brunsonia benewah
.
Fig. 182. Anterior gonopods, posterior view. Fig. 183. Tip of left anterior gonopod, posterior view. Fig. 184. Left posterior gonopod coxite, posterior view. Figs. 185–189. Male legs 3–7, respectively.
Etymology
: The O’Hara Campground is located on the Selway River, hence the adjectival species epithet.
Description
:
Male
holotype
: Length, 10.0 mm. Twenty ocelli in triangular eyepatch. Metazonites with low shoulders on all trunk rings from three to 25. Color light brown, lightly marked darker purplish brown, markings more distinct anteriorly. Legpairs one and two reduced, three to seven enlarged, mesal knobs present only on femora four and five, mesal on both (
Figs. 175–179
). Anterior gonopods (
Fig. 173
,
190
) simple, curved, with median basal process well-developed. Posterior gonopod coxites (
Figs. 181
,
191
) complex, with short, lateral retrorse process, mesal fimbriate region very much reduced; subterminal process, curved, thin, acute, terminal process with two lamellate divisions reflexed. Legpair 10 coxae of normal size, with small glands, legpair 11 femora with long, thin, dorsally directed knobs.
Female
10 mm
long, similar to male in nonsexual characters.
Distribution
: This species has been collected only at the
type
locality.
Notes
: Lack of material prevented SEM examination. The posterior gonopod coxites are unique among
Brunsonia
species and it is difficult to homologize the processes with the standard pattern, as set by
B. albertana
.
However, the process labeled
a
in
Fig. 181
may actually be that process. The channeled process seems obsolete, but there is a small pore subapically (not shown in the drawings but observed later).