A new species, Hoya honglenae and the first report of H. acuminata and H. phuwuaensis (Apocynaceae, Asclepiadoideae) in the flora of Vietnam
Author
Nguyen, Van Canh
0000-0001-9578-0342
Institute of Applied Technology, Thu Dau Mot University, No. 6 Tran Van On Street, Phu Hoa Ward, Thu Dau Mot City, Binh Duong Province, Vietnam.
nguyenvancanh@tdmu.edu.vn
Author
Averyanov, Leonid V.
0000-0001-8031-2925
Komarov Botanical Institute of the Russian Academy of Sciences, Prof. Popov Street 2, 197376, St. Petersburg, Russia.
av_leonid@mail.ru
Author
Nguyen, Quoc Bao
0000-0001-8172-8578
Institute of Tropical Biology Vietnam Academy of Science and Technology, 85 Tran Quoc Toan, District 3, Ho Chi Minh City, Vietnam.
quocbao.nguyen98@gmail.com
Author
Dang, Van Son
0000-0001-8681-4141
Institute of Tropical Biology Vietnam Academy of Science and Technology, 85 Tran Quoc Toan, District 3, Ho Chi Minh City, Vietnam.
dvsonitb@gmail.com
Author
Nguyen, Van Khuong
0000-0002-1264-5178
Institute of Applied Technology, Thu Dau Mot University, No. 6 Tran Van On Street, Phu Hoa Ward, Thu Dau Mot City, Binh Duong Province, Vietnam.
nguyenkhuongbanme@gmail.com
Author
Maisak, Tatiana V.
0000-0001-5919-6755
Komarov Botanical Institute of the Russian Academy of Sciences, Prof. Popov Street 2, 197376, St. Petersburg, Russia.
tmaisak@mail.ru
Author
Truong, Ba Vuong
0000-0003-3452-8455
Institute of Tropical Biology Vietnam Academy of Science and Technology, 85 Tran Quoc Toan, District 3, Ho Chi Minh City, Vietnam.
bavuong2019@yahoo.com
text
Phytotaxa
2023
2023-12-27
632
1
60
68
https://phytotaxa.mapress.com/pt/article/download/phytotaxa.632.1.5/51389
journal article
10.11646/phytotaxa.632.1.5
1179-3163
10435180
Hoya phuwuaensis
Kidyoo (2016: 218)
.
H.
sect.
Peltostemma
Schlechter (1916: 14)
(
Fig. 3
).
Type
:—
THAILAND
.
Phu Wua Wildlife Sanctuary
,
Bueng Kan Province
,
200 m
, 2008,
M. Kidyoo
1035
(
holotype
BKF
,
isotype
BCU
, not seen, fide
Kidyoo 2016
)
.
Description
(based on specimens from
Vietnam
):—Slender creeping epiphytic herbaceous vine to
2 m
long with pure white latex in all vegetative parts.
Stem
terete,
2–2.5 mm
in diameter, pale gray brown, scabrous; internodes (2.5)3–6(7) cm long.
Leaves
shortly petiolate; petiole gray brown, scabrous, (3.5)4–7(8) mm long, (1.8)2–2.2(2.5) mm in diameter; leaf blade thick, coriaceous, elliptic, acute, base cuneate, scabrous, (5.5)6–9(11) cm long, (1.8)2– 3(3.2) cm wide, margins entire to slightly undulate, wrinkled, or revolute; adaxial surface dark grassy dark green with large, irregular white blotches; abaxial surface uniformly light pale green; median and lateral veins almost invisible.
Inflorescence
extra-axillary, pendulous, 20–28-flowered umbelliform raceme; peduncle slender, dirty purple brown, (4.5)5–5.5(6) cm long, scabrous; rachis perennial, fleshy.
Pedicels
of the peripheral flowers
1.4–1.5 cm
long, light pinkish to almost white.
Calyx
persistent, deeply 5-lobed, lobes white to pinkish, narrowly ovate, obtuse,
1.4–1.6 mm
long, scabrous outside.
Corolla
discoid, with 5 broadly triangular lobes, white,
0.8–1.2 cm
in diameter; adaxial surface densely pubescent with long straight hairs along the margins of the lobes; abaxial surface glabrous; lobes reflexed, distally revolute,
3.2–4 mm
long, apex acute.
Corona
purple red,
4.8–5 mm
in diameter; outer angle of corona lobe narrowly ovoid,
2–2.2 mm
long, almost straight or distally slightly up-cured, with round apex, upper surface convex; inner angle of corona lobes erect, narrowly conoid, acute,
1.3–1.4 mm
tall, much higher than outer angles.
Anther appendages
yellow with a white apical part, erect, narrowly triangular, acute,
0.8–1 mm
long, exceeding the apex of inner corona angles.
Pollinarium
about
0.3 mm
long; pollinia pale yellow, obliquely oblong,
0.3–0.31 mm
long, narrowly winged, truncate, margins pellucid; caudicles short, hyaline, with broad wing, light pale yellow; translators, pale yellow, obscurely triangular; corpusculum brown, obtriangular, laterally compressed.
Pistil
dull pink, about
1 mm
long, of two separate, glabrous, wine bottle shaped ovaries.
Stigma head
white, broadly conoid,
0.7–0.8 mm
tall and wide.
Fruits
not seen.
FIGURE 3.
Hoya phuwuaensis
Kidyoo.
A, B.
Flattened
portion of plant with flowers (A) and leaves (B).
C.
Latex on the stem cut.
D.
Inflorescence, view from above.
E.
Inflorescence, frontal view.
F.
Flowers, side view.
G.
Pedicel and flowers, frontal view and view from back.
H.
Corolla, adaxial side.
I.
Corona, view from above and from below.
J.
Corona, side view.
K.
Flower, sagittal section.
L.
Pollinarium, view from different sides.
M.
Calyx, with exposed pistil. All photos made from plant used for the preparation of the herbarium specimen
BV 1667
by Nguyen Van Canh, photo correction and design by L. Averyanov and T. Maisak.
Habitat and phenology
:—Epiphyte on tall trees in broad-leaved evergreen mountain forests, commonly along stream valleys, often on
Ficus
sp.
Flowers in May.
Distribution
:—
Vietnam
,
Dak Lak Province
(Lak District, Chu Yang Sin Mountains). NE
Thailand
(
Bueng Kan Province
, Phu Wua Wildlife Sanctuary).
Conservation status:
—In
Vietnam
, only one small population of this rare species has been discovered until now. It was found in an area affected by occasional deforestation, seriously impacting habitat quality. The species may also be threatened in the future by commercial collecting. Additionally, a very small area of the occurrence and occupation identifies conservation species status as Nationally Critically Endangered (nationally CR), following the formal criteria proposed by UIUCN (2022) as follows:A3c,d; B1a,b(i–iii,v)+2a,b(i–iii,v); C1+2(i,ii); D1;—population reduction suspected to be met in the future (A3), a decline in area of occupancy (AOO), extent of occurrence (EOO), and habitat quality (c), and actual or potential levels of exploitation (d);—the area of occurrence <
100 km
² (B1) and the area of occupancy <
10 km
² (B2), with 1 known population (a) and continuing observed decline of extent of occurrence (bi); the area of occupancy (bii); area, quality of habitat (biii); the number of mature individuals (bv);—the number of mature individuals <250, estimated or projected continuing decline in 25% in 3 years or 1 generation (C1), and observed, estimated, projected or inferred continuing decline when the number of mature individuals in each subpopulation (C2(i)) ≤ 50 and % of mature individuals in one subpopulation = 90–100% (C2(ii));—the number of mature individuals <50 (D). The discovered population of this species in
Vietnam
is distant from its alone early-known location in NE
Thailand
, more than
800 km
to the SE. According to available information, the species may already be extinct from nature in
Thailand
(Kydoo 2016), and the Vietnamese population represents currently the only natural refugium of this remarkable taxon in nature.
Note
:—The species is easy to cultivate and remarkable for its beautiful flowers and white-green tessellated leaves. The plant has good prospects for cultivation as an ornamental plant.
Studied specimens
:—
Vietnam
,
Dak Lak Province
, Lak District, Chu Yan Sin Mountains, epiphyte on
Ficus
sp.
,
16 September 2021
,
Truong Ba Vuong & Nguyen Van Canh, BV 1667
(VNM 00043091, photos—LE 01124052 https://en.herbariumle.ru/?t=occ&id=165926).
Vietnam
,
Dak Lak Province
, Lak District, herbarium prepared from cultivated plants by Nguyen Van Canh
s.n.
in
May 2022
, det.:
May 2022
,
Nguyen Van Canh, AL 1397
(LE 01169047 https://en.herbariumle.ru/?t=occ&id=140563).