Systematics and phylogeny of Philautus Gistel, 1848 (Anura, Rhacophoridae) in the Western Ghats of India, with descriptions of 12 new species Author Biju, S. D. Author Bossuyt, Franky text Zoological Journal of the Linnean Society 2009 2009-02-28 155 2 374 444 https://academic.oup.com/zoolinnean/article-lookup/doi/10.1111/j.1096-3642.2008.00466.x journal article 10.1111/j.1096-3642.2008.00466.x 0024-4082 5445358 PHILAUTUS OCHLANDRAE GURURAJA ET AL ., 2007 ( FIGS 52A, B , 53A; TABLE 2) Type material: Holotype , ZSI/WGFRS/V/A/632, an adult male, SVL 25.6 mm , collected from Kakkayam Reserve Forest, Calicut District, Kerala , India ; ( Figs 52A, B , 53A ). This combination of characters can also be observed in variations of Philautus akroparallagi sp. nov. ( Fig. 4D ) and P. signatus ( Fig. 55B ). However, P. ochlandrae differs from P. akroparallagi sp. nov. by its medium male adult size, SVL 23.9 ± 1.3 mm , N = 5 (vs. small male adult size, SVL 20.7 ± 1.0 mm, N = 16), rounded snout (vs. pointed), head wider than long, HW 8.3 ± 0.3 mm , HL 6.6 ± 0.6 mm , N = 5 (vs. head length equal to width, HW 7.8 ± 0.4 mm , HL 7.8 ± 0.5 mm , N = 10), and from P. signatus by its rounded snout (vs. pointed), canthus rostralis rounded (vs. acute), head wider than long, HW 8.3 ± 0.3 mm , HL 6.6 ± 0.6 mm , N = 5 (vs. head length subequal to width, HW 10.8 ± 1.1 mm , HL 10.6 ± 1.1 mm , N = 8), and absence of a lingual papilla on the tongue (vs. tongue with a lingual papilla). Figure 51. Holotype of Philautus nerostagona : A, paratype (BNHS 4245); B, holotype (BNHS 4244) from Kalpetta. Description of the holotype : The detailed description and illustrations were published by Gururaja et al. (2007b) . Figure 52. Philautus ochlandrae : A, specimen (SDB 6076) from Kakkayam Reserve Forest; B, specimen (BNHS 4559) from Kakkayam Reserve Forest. paratypes , ZSI/ WGFRS / V /A/633 and ZSI/ WGFRS / V / A/637, two males , and ZSI/ WGFRS / V / A636 , a female, from the holotype locality. Other material studied: SDB 6076 and BNHS 4559, two adult males collected from Kakkayam Reserve Forest , Kerala , India ( Table 2 ) . Diagnosis: Philautus ochlandrae can be distinguished from known congeners by the following combination of characters: (1) medium male adult size (SVL 23.9 ± 1.3 mm , N = 5); (2) canthus rostralis rounded; (3) snout rounded; (4) pupil with striking golden yellow dentition-like marks in life; (5) light brown to creamy yellowish, with two invariable and distinct golden yellow streaks running from snout, along the side of head, to near the vent. Philautus ochlandrae can be differentiated from all other known species of Philautus from the Western Ghats by its two distinct golden yellow streaks, extending from the snout along the side of head to near the vent, and by a pupil with striking golden yellow dentition-like marks in all colour morphs Variation: Measurements of six specimens including the type series are given in Table 2 . Distribution and natural history: Philautus ochlandrae is known only from the type locality in the Kakkayam Reserve Forest of the Calicut District ( Fig. 53A ). The type series was collected from the hollow tube of the internodal region of reed brake, about 2.5 m above the ground. Breeding (amplexus) and oviposition also take place inside hollow reeds ( Gururaja et al. , 2007b ).