Genetic and morphological evidence for two species of Leucocarbo shag (Aves, Pelecaniformes, Phalacrocoracidae) from southern South Island of New Zealand Author Nicolas J. Rawlence Author R. Paul Scofield Author Hamish G. Spencer Author Chris Lalas Author Luke J. Easton Author Alan J. D. Tennyson Author Mark Adams Author Eric Pasquet Author Cody Fraser Author Jonathan M. Waters Author Martyn Kennedy text Zoological Journal of the Linnean Society 2016 177 676 694 journal article 10.1111/zoj.12376 b49e437b-c2a1-4e2a-a683-84c637a0535b 270312 LEUCOCARBO CHALCONOTUS OTAGO SHAG GRACALUS CHALCONOTUS G. R. GRAY, 1845 Diagnosis . A species of Leucocarbo most closely related to L. stewarti and L. onslowi but distinguished from these species by the plumage characters and allometries outlined in Table 1 . Distribution . Formerly the eastern South, NZ . Leucocarbo chalconotus bones have been recorded from Late Quaternary and archaeological deposits along the entire eastern coastline of South (e.g., Worthy, 1998a ; Smith, 2011 ). Now restricted to Otago from Lake Ki-Wainono to The Sisters (based on historical museum skins and modern specimens), with modern vagrants north to Banks Peninsula (see Fig. 1 ). Rare modern beach wrecks on Stewart ( Rawlence et al ., 2015 ).