New Australasian Parathalassiinae (Diptera: Dolichopodidae sensu lato) Author Brooks, Scott E. 0000-0001-7915-1941 Diptera Unit, Canadian National Collection of Insects, Agriculture and Agri-Food Canada, K. W. Neatby Building, 960 Carling Avenue, Ottawa, Ontario, K 1 A 0 C 6, CANADA & scott. brooks @ agr. gc. ca; https: // orcid. org / 0000 - 0001 - 7915 - 1941 scott.brooks@agr.gc.ca Author Cumming, Jeffrey M. 0000-0002-3619-3700 Diptera Unit, Canadian National Collection of Insects, Agriculture and Agri-Food Canada, K. W. Neatby Building, 960 Carling Avenue, Ottawa, Ontario, K 1 A 0 C 6, CANADA & jeff. cumming @ agr. gc. ca; https: // orcid. org / 0000 - 0002 - 3619 - 3700 jeff.cumming@agr.gc.ca text Zootaxa 2022 2022-09-22 5188 6 521 543 journal article 148081 10.11646/zootaxa.5188.6.2 ecb5de27-f5fe-4b16-903b-633548186b0a 1175-5326 7103618 BF01B6B4-3415-41A1-86A0-F4B187541A55 Chimerothalassius Shamshev & Grootaert Chimerothalassius Shamshev & Grootaert, 2002: 131 . Type species: Chimerothalassius ismayi Shamshev & Grootaert, 2002 , by original designation. Diagnosis. The genus Chimerothalassius is distinguished from other parathalassiine genera by the following characters: head with gena scarcely projected below eye ( Figs 12, 14 , 27 ), mouthparts directed ventrally with fleshy labellum ( e.g. , Figs 14 , 27 ), palpus elongate and narrow ( Figs 12, 14 ), abruptly capitate apically and narrow basally ( Figs 19 , 27, 28 ), or broadly subtriangular; thorax with prosternum fused to proepisternum forming precoxal bridge, scutellum with 1 pair of strong dorsally directed setae near apex ( e.g. , Fig. 14 ); legs with fore coxa lacking field of short stout spinose setae on anterior surface, tarsomere 5 of each leg with medial apical projection, although sometimes weakly developed; wing ( Fig. 22 ) with R 1 reaching costa before middle of wing, crossvein bm-m complete or incomplete, cell dm absent without veins M 2 and dm-m, CuA rounded, cell cua convex apically, CuA+CuP absent or vestigial, anal lobe not developed; male terminalia with hypopygium small ( Figs 26, 29, 30 ), right epandrial lamella usually with dorsally directed ventral process ( Figs 17 , 24 , 32 ), cerci symmetrical (or nearly so) and moderately short, hypoproct projected ( Figs 18 , 25 , 33 ); female abdomen with apical segments retracted into segment 5 ( Fig. 21 ), terminalia with syntergite 9+10 undivided and bearing acanthophorous setae and cercus narrowly rounded apically with prominent apical or preapical seta, or syntergite 9+10 divided and bearing acanthophorous spines and cercus pointed apically without apical seta ( Fig. 21 ). Remarks. Species of Chimerothalassius are known from the Caribbean, Costa Rica , New Zealand ( Figs 3–7 ) ( Brooks & Cumming 2018 ) and now New Caledonia ( Figs 8–10 ) ( Cumming & Brooks 2019 , as “Undescribed genus [ New Caledonia ]”). The genus is found on rocky, stony or sandy habitats of coastal beaches ( Figs 4–7 ) and emerged rocks in rivers ( Figs 9, 10 ) ( Shamshev & Grootaert 2002 ; Brooks & Cumming 2018 ; Cumming & Brooks 2019 ).