A review of the New Caledonian Arpactophilus (Hymenoptera: Crabronidae) Author Breitkreuz, Laura C. V. Author Ohl, Michael Author Engel, Michael S. text Zootaxa 2016 4063 1 1 66 journal article 37206 10.11646/zootaxa.4063.1.1 9608b695-6310-4c63-bd94-46bda058e6d8 1175-5326 270412 80401ED8-C6BA-4420-9109-854C5CC1E88D Arpactophilus caac sp. nov. ( Figs. 20 , 35–36 ) Diagnosis. Arpactophilus caac is the only New Caledonian species in the genus with the combination of a black, projecting clypeus with a nearly straight apical margin medially ( Fig. 35 ), yellow mandible, the frontal carina forming a projecting tip on the basal clypeal margin, a yellowish flagellum about as light or only little darker than the scape, a dorsally-interrupted occipital carina ( Fig. 6 ), two submarginal cells in the forewing, yellow legs, a flat metasomal sternum II without a bulge (as in Fig. 1 ), and a dark brown metasoma ( Fig. 36 ). Description. FEMALE: Total length 3.0– 3.7 mm , mean = 3.4 mm ; forewing length 2.2–2.3 mm , mean = 2.2 mm . Body black or brown, with areas of yellow. Yellow: mandible; labrum; palpi; scape; pedicel; flagellum (slightly darker towards apex); anterior pronotal margin; pronotal lobe; tegula; legs. Wings hyaline; pterostigma brown. Head about 0.9 × as long as wide in frontal view. Apical margin of clypeus projecting medially, medioapical margin nearly straight ( Fig. 35 ). Apical margin of labrum bilobed. Ventral mandibular tooth about ¼–1/5 of total mandibular length, not reaching opposite mandibular base. Palpal formula probably 4:3 (not completely visible). Frons imbricate with sparse punctation and long setae. Frontal carina present from median ocellus to upper half of clypeus, forming a distinct point at basal clypeal margin. Scape 3.9 × as long as wide. Ocellar triangle anterior of tangent between upper posterior orbits of compound eyes, lateral ocelli anterior of tangent by approximately their diameter. Occipital carina interrupted dorsally ( Fig. 6 ). Gena imbricate with sparse punctation and long setae, without tubercles, spines, or dorsoventral carina. Hypostomal midventral line carinate, not angulate. FIGURES 29–34. Facial views and lateral habitus of ♀ holotypes of species of Arpactophilus Smith. 29. Arpactophilus arhoe sp. nov. , facial view; 30. A. arhoe , lateral habitus;31. A. brochus sp. nov. , facial view; 32. A. brochus , lateral habitus; 33. A. bwatoo sp. nov. , facial view; 34. A. bwatoo , lateral habitus. Mesosoma about 1.7 × as long as wide in dorsal view. Propodeum about 0.9 × as long as wide in dorsal view. Mesosoma imbricate with sparse punctation ( Fig. 36 ), except posterior part of dorsal surface of propodeum; lateral surface of propodeum also imbricate and transversely carinulate; dorsal surface of propodeum also reticulate medially, lateral and posterior areas of dorsal surface of propodeum transversely carinulate ( Fig. 20 ). Slightly pitted sulcus present posterior to fine mesoscuto-scutellar sulcus. Hypersternal sulcus represented by fine line. Metafemur 3.0 × as long as wide. Metatibia without differently colored area apically. Pretarsal claws without teeth. Forewing with two submarginal cells; anterior border of submarginal cell II slightly shorter than posterior border. Hind wing with five distal hamuli. Metasoma polished, sparsely punctured ( Fig. 36 ). Metasomal sternum II without bulge (as in Fig. 1 ). Pygidium with narrow row of silk setae apically. MALE: Difference from female aside from genitalic structures: apical half of clypeus yellow. Metasomal sternum VIII neither broadened nor narrowed. Remarks. Arpactophilus caac is somewhat similar to A. pwaamei , A. kraussi , and A. tayo in the head shape, size and general coloration. Arpactophilus kraussi has a finer sculpture on the propodeal dorsum. Arpactophilus caac can be distinguished from A. tayo easily by the black coxae of the latter. The coarse macro-punctation on the frons of A. pwaamei distinguishes it from A. caac with its fine punctation. Material examined. HOLOTYPE ♀: “ NEW CALEDONIA : 9.2 km NE Col d’Amieu on Rte. 5 slopes of Mt Rembai, 375–675 m , 23.IX.1979 ; W.C. Gagne, G.M. Nishida, Colls. BISHOP Museum Acc. No. 1979.380 [ BPBM ]. PARATYPES 7♀, 1♂ : ♀: ( 1x ) NEW CALEDONIA 8905: 20°58’S x 165°17’E , 500m , Pic d’Amoa, N slopes. 24 Nov 2001 – 31 Jan 2002 . Burwell, Monteith. Malaise [QM]; ( 1x ) NEW CALEDONIA 8926: 21°11’S x 165°18’E , 850m , Aoupinie top camp. 23 Nov 2001 – 1 Feb 2002 . Burwell, Monteith. Malaise [QM]; ( 1x ) NEW CALEDONIA 9964: 20°33’S x 164°46’E , 900m , Mt Panie track. 25 Sep–19 Nov 2000 . Skevington & Burwell. Malaise [ ZMB ]; ( 1x ) “Nouvelle Calédonie, Pinda 30m , 18.IV.1995 , Réc. Chazeau & Jourdan [ CAS ]; ( 2x ) “Nouvelle Calédonie, Mont Koghis. 350m , 25.IV.1995 , Réc. Chazeau & Jourdan [ MNHN ]; ( 1x ) “ NEW CALEDONIA Col d’Amieu, 650m 21.III.1968 ; T. C. Maa, Collector, BISHOP [ BPBM ]. ♂: “ NEW CALEDONIA Mont Panie, 700 m , 27-29.VII.1971 ; J.L. Gressitt, Collector, BISHOP museum [ BPBM ]. Etymology. The specific epithet is taken from the name Caac , one of the New Caledonian native languages. It is treated as a noun in apposition.