A revision of the genera Pseudosmittia Edwards, 1932, Allocladius Kieffer, 1913, and Hydrosmittia gen. n. (Diptera: Chironomidae, Orthocladiinae) 2849 Author Ferrington Jr, Leonard C. Author Saether, Ole A. text Zootaxa 2011 2011-04-29 2849 1 1 314 https://biotaxa.org/Zootaxa/article/view/zootaxa.2849.1.1 journal article 10.11646/zootaxa.2849.1.1 1175­5334 5288200 Pseudosmittia triangula (Tokunaga) ( Fig. 74 C–G ) Smittia triangula Tokunaga, 1964: 524 . Pseudosmittia triangula (Tokunaga) ; Cranston and Martin (1989: 262) ; Saether (2004: 296). Material examined : PALAU (BELAU): Koror Island , at light, holotype male, 2.v.1957 , C. W. Sabrosky ( USNM 66540 ); 2males same data as holotype, except 29.iv.1957 ; 1 female , same data as holotype , except ix.1952 , N. L. H. Krauss ( BBM ) ; Malakal Island , allotype female, 2.v.1957 , C. W. Sabrosky ( USNM ) ; Sonsorol Island , 1 male , 13.ix.1952 , N. L. H. Krauss ( BBM ) , Babelthuap Island , E . Ngatpang , light trap , 1 male , 9.xii.1952 , J. L. Gressitt ( BBM ) . SEYCHELLES : Silhouette , Grande Barbe , 4 males , 6.i.222, J. Gerlach ; Silhouette , La Passe (above Dauban mausoleum), Malaise , 1 male , 1–4.vii.2000 , J. Gerlach ( ZMBN ) . CHINA : Jiangxi Prov. , Yifeng Co. , Gongshan , Xihe , 1 male , 8.vi.2004 , C.C. Yan ( XW ) . Diagnostic characters : The male imago can be separated from the other members of the group by lacking an apical antennal seta, an AR of about 0.6, and 10–14 dorsocentrals. The female imago is characterized by an AR of 0.32–0.37, 5 sensilla clavata at the apex of third palpomere and 8 dorsocentrals. Male imago (n = 3–4, except when otherwise stated) Total length 1.27–1.35, 1.32 mm . Wing length 0.68–0.75, 0.73 mm . Total length/wing length 1.78–1.86, 1.82. Wing length/length of profemur 2.53–2.61, 2.57. Coloration dark brown. Head ( Fig. 74 D ). AR 0.62–0.64. Terminal flagellomere 1.71–1.78 µm long, no apical strong seta. Temporal setae 5–6, 5, consisting of 2–3, 2 inner verticals, and 3–4, 3 outer verticals, no postorbitals. Clypeus with 6–9 setae. Tentorium 89–94, 91 µm long, 18–23, 21 µm wide. Stipes not measurable. Palpomere lengths (in µm): 18–21, 20; 25–32, 28; 43–46; 50–53; 75–90. Third palpomere with 2 sensilla clavata. Thorax. Antepronotal lobes reduced medially, antepronotal lateral setae not observable. Dorsocentrals 10–14, 13, acrostichals 2, prealars 3–4, 3; no supraalar seta. Scutellum with 5–6, 6 setae. Wing ( Fig. 74 C ; Saether 2004 fig. 29). VR 1.66–1.74, 1.70. VR 2 1.25–1.39, 1.34. Anal lobe weak. Costal extension 11–16 µm long. R 4+5 ending well proximal to apex of M 3+4 ; Cu 1 slightly curved. Brachiolum with 1 seta, other veins bare. Legs. Spur of front tibia 25–41, 35 µm long, of middle tibia 21–27 µm long (only one present), spurs of hind tibia 27–41 µm and 1–23 µm long. Width at apex of front tibia 18–22, 19 µm, of middle tibia 21–24 µm, of hind tibia 27–32 µm. Comb of 11–12 setae, 18–37 µm long. Lengths and proportions of legs as in Table 45 . TABLE 45. Lengths (in µm) and proportions of legs of Pseudosmittia triangula (Tokunaga) , male (n = 3–4. 1–2 for tarsi).

fe	ti	ta1	ta2	ta3	ta4	ta5	LR	BV	SV	BR
p1	270–288, 287	279–315, 234	176–183	77–87	50–54	30–36	32	0.56–0.58	3.93–3.95	3.30–3.44	3.2
p2	288–327	293–329	174	82	59	34	34	0.53	3.95	3.78	–
p3	279–343	302–341	198–210	99–101	99–101	41–45	36–37	0.59–0.62	3.00–3.20	3.23–3.25	4.0

Hypopygium ( Fig. 74 F, G ; Tokunaga 1964 fig. 6f; Saether 2004 fig. 30). Anal point 30–43, 35 µm long, triangular and without apical projection; tergite IX (including anal point) with 10–12 (2) setae; laterosternite IX with 4–5, 4 setae. Phallapodeme 48–62 µm long; transverse sternapodeme 37–43 µm long, no oral projections. Virga ( Fig. 74 E ) 17–25 µm long. Gonocoxite 110–126, 116 µm long; inferior volsella long, reaching to 0.84–0.91, 0.87 gonocoxite length. Gonostylus 65–71, 69 µm long, attenuate; megaseta 5–7, 6 µm long. HR 1.67–1.77, 1.70, HV 1.86–1.96, 1.93. Female imago (n = 1–2) Total length 1.08–1.34. Wing length 0.59–0.63. Total length/wing length 1.85–2.11. Wing length/length of profemur 3.07. Coloration yellow with dark brown vittae and markings. Head. AR 0.32–0.37. Flagellomere lengths (in µm): 48–57, 34–43, 37–41, 37–43, 55–59. Apical seta 32 µm long. Temporal setae 5–6, consisting of 2 inner verticals, and 3–4 outer verticals, no postorbitals. Clypeus with 6–8 setae. Tentorium 85 µm long, 7 µm wide. Stipes 69–78 µm long. Palpomere lengths (in µm): 18–21, 23–25, 43–45, 53–57, 66–69. Third palpomere with 5 sensilla clavata. Coronal suture absent. Thorax. Median antepronotal lobes reduced, antepronotum with 2 lateral setae. Dorsocentrals 8, acrostichals 2, prealars 3–4. Scutellum with 4–6 setae. Wing. VR 1.70–1.76. VR 2 1.30. Anal lobe weak, costal extension 69–80 µm long. R 4+5 ending proximal to apex of M 3+4 ; Cu 1 straight. Brachiolum with 1seta, costa between FR and apex of R 4+5 with 19–21 non-marginal setae, costal extension with 3–7 non-marginal setae, R with 5–8 setae, R 1 with 1–2, R 4+5 with 5–10, setae, other veins bare. Legs. Spur of front tibia 23–25 µm long, spur of middle tibia 24–25 µm long (only 1 present), of hind tibia 34 µm and 21 µm long. Width at apex of front and middle tibiae each 18–21µm, of hind tibia 26–31 µm. Comb of 11– 12 setae, 21–34 µm long. Lengths and proportions of legs as in Table 46 . TABLE 46. Lengths (in µm) and proportions of legs of Pseudosmittia triangula (Tokunaga) , female (n = 1–2).

fe	ti	ta1	ta2	ta3	ta4	ta5	LR	BV	SV	BR
p1	198–207	234–237	122–126	41–45	36	27	27	0.51–0.54	4.20–4.28	3.50–3.59	2.2–2.4
p2	270–279	297–302	135–140	59–63	45	27	27–32	0.45–0.46	4.33–4.44	4.16–4.20	2.4–2.7
p3	243–252	270–275	153	72	72	27–32	27–32	0.56–0.57	3.41	3.41	2.4–3.0

Abdomen. Tergite I bare, T II–VII each with 8–10 setae, T VIII with 16 setae. Sternites I and II bare, S III–V each with 2–4 setae, S VI and VII each with 7, S VIII with 25 setae. Genitalia. Gonocoxite with 2–3 strong and 2 weak setae. Tergite IX divided with altogether about 6 setae. Cercus 48 µm long. Seminal capsule 64–73 µm long, 41–50 µm wide, ovoid, dark. Notum 55–57 µm long. Pupa and larva Unknown. Distribution The species is known from Palau , the Marshall Islands , the Seychelles and the Jiangshi Province in China .