Resolving taxonomic issues of cicadas (Hemiptera: Cicadidae) including new combinations, new synonymies, and revised status, with updates on the diversity of the Brazilian cicada fauna and new records for four South American countries
Author
Sanborn, Allen F.
text
Zootaxa
2023
2023-07-20
5318
3
339
362
http://dx.doi.org/10.11646/zootaxa.5318.3.2
journal article
57485
10.11646/zootaxa.5318.3.2
39e59e73-ea57-4c57-965f-06bd576d5739
1175-5326
8166896
D7216A8E-B6C2-4A34-9EC6-CACC3D9951E3
NEW COMBINATIONS AND SYNONYMIES IN
ZAMMARINI
BASED ON
GOEMANS (2016)
Goemans (2016)
studied the
Zammarini
providing evidence for numerous taxonomic changes. The taxonomical acts within the dissertation were not considered to be valid for nomenclatural purposes following Article 8 of the
International Code of Zoological Nomenclature
(ICZN 1999) and were disclaimed for such purposes in that work (
Goemans 2016
). The specifics and long history of confusion for the application of
Zammarini
taxa is described in
Goemans (2016)
. He was able to study almost all
type
specimens and clarify the taxonomy of the group as a whole. The original work (
Goemans 2016
) should be consulted for summary information on the morphology, history, and distribution of each species studied.
Unfortunately, he seems to have moved on to a different area of study and has yet to publish the results formally. Some of his findings have already been applied to update the taxonomy of numerous species (e.g.,
Sanborn 2019a
;
2020c
) and other changes are being reported in the literature even though the changes are unofficial under the
Code
(e.g.,
Nunes
et al
. 2023
). Most of his remaining nomenclatural data is presented here to reassign species to clarify the taxonomy of the
Zammarini
as his work is important in correcting historic misconceptions and the unofficial results are being incorporated into the more recent literature. The new combinations are provided here to make the changes suggested by
Goemans (2016)
official under the
Code
so that correct taxonomy can be applied to these taxa in the future.
With respect to Brazilian species,
Zammara eximia
Erichson, 1848
was determined to possess three tarsomeres and is reassigned to
Zammaralna eximia
(
Erichson, 1848
)
n. comb.
, as presented in the species list of
Nunes
et al
. (2023)
. Similarly,
Zammara strepens
Amyot & Audinet-Serville, 1843
was determined to have three tarsomeses and a narrow head so it is reassigned to
Orellana strepens
(
Amyot & Audinet-Serville, 1843
)
n. comb.
, also as presented in the species list of
Nunes
et al
. (2023)
. In addition, it was determined that the images and distribution of
Orellana pollyae
Sanborn, 2011a
were the same as
O. strepens
n. comb.
so that
Orellana pollyae
Sanborn, 2011a
n
. syn.
becomes a junior synonym of
Orellana strepens
(
Amyot & Audinet-Serville, 1843
)
n. comb.
Nunes
et al
. (2023)
included
O
.
pollyae
n. syn.
in their list of species as a junior synonym of
O. strepens
n. comb.
but did not make the change official under the
Code
.
There are additional taxonomic changes reported in
Goemans (2016)
in species of
Zammarini
not inhabiting
Brazil
.
Orellana brunneipennis
Goding, 1925
and
Orellana pulla
Goding, 1925
were shown to possess two tarsi instead of three so they are reassigned to become
Zammara brunneipennis
(
Goding, 1925
)
n. comb.
and
Zammara pulla
(
Goding, 1925
)
n. comb.
, respectively.
Zammara brunneipennis
(
Goding, 1925
)
n. comb.
,
n. syn.
,
Zammara pulla
(
Goding, 1925
)
n. comb.
,
n. syn.
, and
Zammara erna
Schmidt, 1919
n. syn.
are junior synonyms of
Zammara intricata
Walker, 1850
. Finally,
Zammara medialinea
Sanborn, 2004
n. syn.
is a junior synonym of
Zammara luculenta
Distant, 1883
.
Goemans (2016)
illustrated the variability in coloration and wing patterns in these species which led to the description of multiple taxa for the highly variable species involved.