Phylogeny, taxonomy and flower-breeding ecology of the Colocasiomyia cristata species group (Diptera: Drosophilidae), with descriptions of ten new species Author Takano, Kohei Takenaka Author Gao, Jian-Jun Author Hu, Yao-Guang Author Li, Nan-Nan Author Yafuso, Masako Author Suwito, Awit Author Repin, Rimi Author Pungga, Runi Anak Sylvester Author Meleng, Paulus Ak Author Kaliang, Clement Het Author Chong, Lucy Author Toda, Masanori J. text Zootaxa 2021 2021-12-08 5079 1 1 70 journal article 2931 10.11646/zootaxa.5079.1.1 010695ef-a142-42ad-81d7-2e7ed1f7e89e 1175-5326 5766402 2B118B81-7353-4A2C-A892-5466DFC83230 2.2. Colocasiomyia cristata species complex, new Diagnosis . Mid tibia with only 1 large, apical seta (ch.15-1; Fig. 7D ). Male abdominal sternite VI with a pair of processes bearing prominent seta on apex and stout setae on submedial to distal portion (ch.26,27-1; Fig. 9G–K ). Lateral pieces of subepandrial sclerite broad plates somewhat sclerotized at least on ventral margin (ch.37-4; Fig. 10D ). Phallal sheath thicker than wide (ch.43-0; Fig. 11H,I ). Shared characters. Anterior reclinate orbital seta small, situated behind proclinate orbital seta; additional, interfrontal setulae on fronto-orbital plate present (ch.2-0; Fig. 3F ). Eye with stout, dense interfacetal setulae (ch.3- 1; Fig. 3H ). Distance between antennal sockets narrower than half of socket width (ch.4-1; Fig. 3G ). Cibarial, medial sensilla in parallel rows narrower than sensilla campaniformia (ch.5-2; Fig. 4B ). Neither additional dorsocentral setae on presutural area nor prescutellar acrostichal setae present; acrostichal setulae in 4 rows (ch.8-1, 9-0; Fig. 5C ). Katepisternal setae small, shorter than postpronotal setae (ch.10-1; Fig. 5F ). Basal scutellar setae long; posterior tip beyond half of apical scutellar setae (ch.11-1; Fig. 5C ). Apical scutellar setae nearer to each other than to basal scutellar seta ( Fig. 5C ). Costal setae in middle row all weak, trichoid (ch.12-2; Fig. 6C ). Patch covered with only minute pubescence absent on anterodorsal portion of fore tarsomere I (ch.13-1; Fig. 7B ). Male abdominal sternite V medially concaved on posterior margin. Female abdominal sternites III and IV longer than wide (ch.19,20-1). Male abdominal sternite VI not pubescent at least partly (ch.23-0; Fig. 9G–K ). Epandrial phragma small (narrower than epandrial anteroventral corner) expansion on antero-subapical margin (ch.33-2; Fig. 10C ). Surstylus vestigial, very narrow process, apically with 1 or 2 tiny, trichoid seta(e) (ch.34,35-1; Fig. 10E ). Median piece of subepandrial sclerite broad, somewhat quadrate plate (ch.36-0). Cercal, ventral/apical portion more or less differentiated from upper portion (ch.38-0; Fig. 10C ). Phallal sheath not pubescent, gently curved ventrad, projected anteriad at dorsobasal corner in lateral view, apically not bilobed (ch.40-1, 41-3, 42-0, 44-1; Fig. 11H ). Gonocoxites long, narrow slips ( Fig. 11H ). Epiproct with pubescence ( Fig. 12C ). Included species. Colocasiomyia cristata , C. sulawesiana , C. kinabaluana sp. nov. , C. kotana sp. nov. and C . matthewsi sp. nov.