Pamphagidae (Orthoptera: Caelifera) of North Africa: key to genera and the annotated check-list of species
Author
Massa, Bruno
text
Zootaxa
2013
3700
3
435
475
journal article
10.11646/zootaxa.3700.3.7
371f30c1-ff9d-4fde-9429-fc9d284caf03
1175-5326
249376
E3FA2E4B-DDF4-4FDB-8CF1-FFBA215D40CA
Tmethis
Fieber, 1853
(type-species:
Gryllus
cisti
Fabricius)
Fieber (1853) did not explain the origin of the name
Tmethis
; thank to my Greek colleague Haralabos Tsolakis, it was possible to trace a link with the verb "temno" (τέΜνω), which means "to cut", possibly referred to the particular shape of pronotum, with a distinct sulcus between prozona and metazona. I consider that
Tmethis
is of masculine gender, and consequently I will agree taxa with it. In his excellent revision of all Palaearctic
Thrinchinae
, Uvarov (1943) pointed out the high variability of many species, mainly belonging to genera
Tmethis
and
Asiotmethis
Uvarov
, that may have wrinkled or smooth integuments; sculpturing of exposed parts (mainly pronotum) may be devoid of the usual tubercles and keel. Thus, taxa belonging to the genus
Tmethis
are much variable, not only in their general colour and in the length of wings, but also in their integuments and body structure, which may be more or less stout or slender. Some specimens appear squat and broad (
Figs. 73, 75
). In addition, the colour of hind tibiae varies from red to yellow and to purple or bluish-violet; it seems that this variation characterizes single populations. The arrangement of species within the genus was much unclear, in particular the separation of
T. cisti
from
T. pulchripennis
. Reading Chopard (1943) it is evident that many populations of
T. pulchripennis
were actually
T. cisti
(e.g., those living on
Djerba
and Kerkenna Is.). The first author who defined characters to identify species was Uvarov (1943); he recognized
T. maroccanus
from
T. cisti
and
T. pulchripennis
by the shape of the metazona of pronotum, which is broadly parabolic with convex margins in the former and triangular, with the sides straight or somewhat incurved in the other two species. Additionally, he identified
T. cisti
and
T. pulchripennis
by the shape of their male epiphallus, and by small differences on the lower valve of the ovipositor and length of spurs of hind tibiae. Thus, records of species in North African countries may be considered reliable only if criteria used by Uvarov (1943) were followed. However, La Greca (1969) considered much variable the shape of lower valve of the ovipositor (according to Uvarov (1943) in
T. pulchripennis
with a distinct shoulder, which appears as a tooth in lateral view, while in
T. cisti
it appears without a projection, not dentate), and identified some
T. pulchripennis
from
Libya
, following criteria of Chopard (1943), based on the pronotum shape, that during this study resulted to be a very variable character. Tegmina length is also a variable character, as supposed by Jannone (1938). Eight subspecies or varieties have been described for
T. cisti
, three for
T. pulchripennis
and three for
T. maroccanus
, in some cases overlapping. I will examine each of them to establish if their taxonomic status is justified.