Diversity and phylogeny of the extinct wasp subfamily Lancepyrinae (Hymenoptera, Bethylidae) revealed by mid-Cretaceous Burmese amber Author Brazidec, Manuel https://orcid.org/0000-0002-0860-8972 Univ. Rennes, CNRS, Geosciences Rennes, UMR 6118, 35000 Rennes, France & State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology and Center for Excellence in Life and Paleoenvironment, Chinese Academy of Sciences, 39 East Beijing Road, Nanjing 210008, China manuel.brazidec@gmail.com Author Legendre, Frederic https://orcid.org/0000-0001-5900-8048 Institut de Systematique, Evolution, Biodiversite (ISYEB), Museum national d'Histoire naturelle, CNRS, Sorbonne Universite, EPHE, Universite des Antilles, CP 50, 57, rue Cuvier, 75005 Paris, France Author Perrichot, Vincent https://orcid.org/0000-0002-7973-0430 Univ. Rennes, CNRS, Geosciences Rennes, UMR 6118, 35000 Rennes, France text Arthropod Systematics & amp; Phylogeny 2023 2023-04-04 81 345 369 http://dx.doi.org/10.3897/asp.81.e96737 journal article http://dx.doi.org/10.3897/asp.81.e96737 1864-8312-81-345 0C2444B0ECFB4A22ACF6F2ACEC0DDCDF B69B5606C9B358B9A9027E3E43F2B66D Gwesped groehni sp. nov. Figs 3 , 4 Etymology. The specific epithet is a patronym honouring Carsten Groehn , who generously made the specimen available for study. The specific epithet is to be treated as a noun in the genitive case. Material studied. Holotype GPIH.5068, a complete female; housed in the Carsten Groehn amber collection (under CCGG no. 11341) of the Leibniz Institute for the Analysis of Biodiversity Change , Hamburg , Germany (GPIH). Type locality and horizon. Noije Bum, Hukawng Valley, Kachin State, northern Myanmar; late Albian-early Cenomanian, ca. 99 Ma, mid-Cretaceous. Diagnosis. As for genus. Description. Body depressed and poorly pubescent (length 3.82 mm). - Head prognathous, quadrate, about as long as wide, with metallic reflections; LH: 0.65 mm, WH: 0.56 mm; WF: 0.39 mm, HE: ca. 0.19 mm; frons flat, slightly punctate; compound eye rounded, located anteriorly on head; clypeus with median lobe only slightly projecting forward, lateral lobe poorly developed; antenna short, barely reaching pronotum posteriorly; scape 2.2 times as long as pedicel (length 0.20 mm); pedicel longer than flagellomeres 1-9; 10 flagellomeres, all compact, as long as wide; flagellomere tapering at apex, flagellomere 10 slightly longer than flagellomeres 1-9 (0.05 mm vs. 0.11 mm); occipital carina present. - Mesosoma narrow with dorsum smooth (length 1.56 mm); propleuron elongate, 'neck-shaped' , visible dorsally; dorsal pronotal area narrower anteriorly, lateral margin slightly incurved, posterior margin straight; anteromesoscutum short, notaulus and parapsidal signum present; mesoscuto-mesoscutellar sulcus conspicuous, widely connecting lateral grooves; mesoscutellum subquadrate; metanotum developed, overlapping mesoscutellum posteriorly; metapectal-propodeal complex with metapostnotal median carina, lateral margin slightly outcurved, posterior corner with small lateral dentiform projection. Fore wing hyaline, posteriorly reaching fourth metasomal segment (LFW = 2.19 mm); C, Sc+R, M+Cu, 1A tubular; 1Rs and 1M aligned; cu-a slightly post-furcal to 1M; Rs+M tubular, reaching apex of pterostigma; pterostigma rounded; 2r-rs arising from distal half of pterostigma; 2r-rs&Rs tubular, reaching wing margin; [2R1] cell large; post-stigmal abscissa of R1 tubular for half-length of [2R1] cell. Legs slender; tibial spur configuration 1-2-2; tarsal claws slightly curved; arolium small. - Metasoma fusiform, smooth (length 1.61 mm); petiole relatively long, conspicuous; six tergites visible, convex; short sting exserted. Figure 4. Line drawing of Gwesped groehni gen. et sp. nov. , holotype female number GPIH.5068. Scale bar: 1 mm.