Taxonomic study of the genus Promalactis Meyrick, 1908 (Lepidoptera: Oecophoridae) X. The commotica species-group, with descriptions of twenty-two new species Author Wang, Shuxia 0000-0002-9316-6661 College of Life Sciences, Nankai University, Tianjin 300071, China & shxwang @ nankai. edu. cn; https: // orcid. org / 0000 - 0002 - 9316 - 6661 shxwang@nankai.edu.cn Author Liu, Chen 0000-0002-9111-6644 College of Life Sciences, Nankai University, Tianjin 300071, China & 348143302 @ qq. com; https: // orcid. org / 0000 - 0002 - 9111 - 6644 348143302@qq.com text Zootaxa 2021 2021-06-01 4980 2 293 330 journal article 6009 10.11646/zootaxa.4980.2.4 88b4aa3d-deff-4fb9-a9eb-1aa9e2726140 1175-5326 4889068 6E352CB3-CF9B-4EE8-B17C-67C88B35FE47 Promalactis motuoensis Wang , sp. nov. ( Figs 12 , 34 , 50 ) Type material. CHINA , Tibet : Holotype ♂ , 80K, Motuo County ( 29.66°N , 95.49°E ), 2059 m , 8.VIII.2017 , leg. MJ Qi & XF Yang , slide No. JYY17672 . Paratypes ( 6♀ ): 4♀ , same data as holotype, except 2059‒2089 m , 6‒ 19.VIII.2017 , slide No. JYY17856 ; 2♀ , same data as holotype, except 2076 m , 8‒9.VIII.2019 , leg. MJ Qi , other same data as holotype , slide No. LC 19323 . Diagnosis. The new species can be distinguished from its congeners by the left valva having an ovate dorsoapical process bearing long dense spines, and the aedeagus triangularly produced and serrate on the margins distally. Description. Adult ( Fig. 12 ). Forewing length 6.0‒7.0 mm. Head: Vertex shining white, frons silvery grey, occiput rust brown. Labial palpus with second palpomere pale rust brown on outer surface, orange yellow on inner surface; third palpomere dark brown except white distally. Antenna with scape silvery white; flagellum black, annulated with white on dorsal surface. Thorax: Dorsum of mesothorax and tegula rust brown. Forewing rust yellow; markings white edged with black scales: costal spot very large, semi-ovate, extending downward crossing anterior angle of cell; basal streak oblique inward to base of fold; antemedian streak from dorsal 2/5 to anterior margin of cell, arched outward, subparallel with basal streak; dorsal streak from beyond basal 2/3 oblique outward to distal 1/5 of posterior margin of cell; tornal spot deep silvery grey with black scales, diffused inward to end of dorsal streak, before tornal spot set a white dot formed by a few white scales; apical spot sub-ovate; terminal spot right below apical spot, separated by black scales; fringe orange yellow except grey around tornus. Hindwing and fringe deep grey. Fore- and midlegs white on dorsal surface, black on ventral surface, tibiae with white scale tufts at apex, mid tibia with a white line at base and middle on ventral side respectively, tarsi white at apex, basal two tarsomeres with wide white distal band; hindleg greyish brown, basal tarsomere white distally, rest tarsomeres greyish white except second and third tarsomeres black basally. Abdomen: Male genitalia ( Fig. 34 ). Uncus with basal 2/5 wide and parallel-sided, thereafter gradually narrowed to rounded apex. Gnathos slightly longer than uncus; mesial plate sub-rounded, with squamous protuberances; basal arms wide basally, uniformly narrowed toward mesial plate distally. Tegumen with lateral arms narrowed anteriorly. Valvae broad sub-rectangular, asymmetrical: apex of left valva rounded, with a large ovate dorsoapical process curved inward and bearing long dense spines, with a small apical spine below dorsoapical process, costa of left valva medially produced to a large sclerotized process bearing dense spines; apex of right valva obtusely rounded, smooth, costa of right valva lined with denser fine spines medially; sacculus wider, setose distally. Saccus almost as long as uncus, slightly narrowed to apex. Juxta short, V-shaped; basal lobe 3/4 length of saccus. Aedeagus straight, almost as long as valva, uniform except slightly widened before apex; apex produced triangularly, serrate. Cornutus absent. Female genitalia ( Fig. 50 ). Apophyses anteriores about 2.5 times length of apophyses posteriores. Lamella postvaginalis heart-shaped, with a triangular notch at middle on posterior margin, arched outward mediolaterally. Antrum slightly narrowed anteriorly, longer than ductus bursae. Ductus bursae membranous, broadened, with a long heavily sclerotized helical belt. Corpus bursae longer than corpus bursae; signum irregularly ovate, bearing dense teeth. Distribution. China ( Tibet ). Etymology. The specific epithet refers to the type locality, Motuo County, Tibet .