Taxonomic study of the genus Promalactis Meyrick, 1908 (Lepidoptera: Oecophoridae) X. The commotica species-group, with descriptions of twenty-two new species Author Wang, Shuxia 0000-0002-9316-6661 College of Life Sciences, Nankai University, Tianjin 300071, China & shxwang @ nankai. edu. cn; https: // orcid. org / 0000 - 0002 - 9316 - 6661 shxwang@nankai.edu.cn Author Liu, Chen 0000-0002-9111-6644 College of Life Sciences, Nankai University, Tianjin 300071, China & 348143302 @ qq. com; https: // orcid. org / 0000 - 0002 - 9111 - 6644 348143302@qq.com text Zootaxa 2021 2021-06-01 4980 2 293 330 journal article 6009 10.11646/zootaxa.4980.2.4 88b4aa3d-deff-4fb9-a9eb-1aa9e2726140 1175-5326 4889068 6E352CB3-CF9B-4EE8-B17C-67C88B35FE47 Promalactis bispina Wang , sp. nov. ( Figs 8 , 30 , 47 ) Type material. CHINA , Yunnan : Holotype ♂ , Taiyanghe National Forest Park ( 22.68°N , 101.03°E ), Pu’er City , 1450 m , 3.VII.2015 , leg. KJ Teng , slide No. JYY17858 . Paratypes ( 5♀ ): same data as holotype, except 4‒5.VII.2015 , slide Nos. JYY17838, JYY17839 . Diagnosis. The new species can be easily distinguished from other members of the group by the male genitalia: the dorsal lobe of the valva has two strong apical spines, the ventral lobe of the valva has a tuft of large setae, and the flagellate juxta has a longitudinal line of dense spines. Description. Adult ( Fig. 8 ). Forewing length 5.0‒6.0 mm. Head: Vertex shining white, frons greyish brown, occiput yellowish brown. Labial palpus with second palpomere pale rust yellow, paler on inner surface; third palpomere blackish brown. Antenna with scape white; flagellum black, with white annulations on dorsal surface. Thorax: Dorsum of mesothorax and tegula rust brown. Forewing orange yellow; markings white edged with blackish brown scales: costal spot sub-triangular, extending obliquely outward crossing anterior angle of cell; basal streak oblique inward to base of fold, straight; antemedian streak from dorsal 2/5 to basal 2/5 of anterior margin of cell, slightly arched outward, parallel with basal streak; dorsal streak from before basal 2/3 extending obliquely outward to below and before posterior angle of cell; tornal spot small, with a blackish grey line extending from beyond it arched upward to end of dorsal streak; apical spot sub-rounded; two small terminal spots interrupt by black scales; fringe orange yellow except greyish brown along distal part of costal margin and around tornus. Hindwing and fringe blackish brown. Fore- and midlegs with ventral surface white; dorsal surface black, tibiae with white spots basally, with white scale tufts at apex, tarsi with basal two tarsomeres white distally; hindleg yellowish white ventrally, blackish brown dorsally, tarsus white except two basal tarsomeres blackish brown distally. Abdomen: Male genitalia ( Fig. 30 ). Uncus triangular, wide at base, narrowed to pointed apex. Gnathos longer than uncus; mesial plate triangular, narrowed to apex, heavily sclerotized; basal arms shorter than mesial plate, narrowly banded. Tegumen sub-trapezoidal, with a shallow anterior emargination. Valva elongate sub-rectangular, length more than 4 times of width; apex shallowly bilobed: dorsal lobe with two strong curved spines apically, ventral spine shorter, ventral lobe very short, truncate at apex, with a tuft of large setae; sacculus wide and folded basally, very narrow distally. Vinculum narrowly banded, rounded on anterior margin. Juxta flagellate, curved, as long as valva, with dense spines along longitudinal line, with an apical spine; basal lobe slender, clubbed. Aedeagus longer than valva, straight, produced to an arrow-like process apically, finely serrate; cornutus absent. Female genitalia ( Fig. 47 ). Papillae anales setose, rounded caudally. Apophyses posteriores 2 times length of apophyses anteriores. Lamella postvaginalis concave at middle on posterior margin, roundly produced posterolaterally. Antrum short, extended to a point posterolaterally. Ductus bursae membranous, narrow basally, broadened distally, with a sclerotized narrow belt bearing 1‒2 short spines. Corpus bursae longer than ductus bursae, granulate; signum longitudinally ovate, large, more than half length of corpus bursae, with two denticles. Distribution. China ( Yunnan ). Etymology. This epithet is from the Latin bis- and spina , referring to the paired apical spines of the valva.