Freshwater Bryozoa of Okinawa, Japan, with descriptions of Rumarcanella gen. nov. (Phylactolaemata: Plumatellidae) and two new species
Author
Hirose, Masato
Author
Mawatari, Shunsuke F.
text
Zootaxa
2011
2732
1
19
journal article
10.5281/zenodo.276554
ec9d7b62-108f-45a6-bb9b-d1c9565c00fa
1175-5326
276554
Plumatella javanica
Kraepelin, 1906
(
Figure 3
)
Plumatella javanica
Kraepelin, 1906
: 143
, figs 1–3;
Annandale 1910
: 42
, 50; 1911: 221;
Toriumi 1952a
: 267
;
Wiebach 1964
: 12
, figs 4–8, pl. 1(4,) pl. 2(5–7), pl. 3(8–12), pl. 4(13); 1967: 77, pls 1, 2(1–8); 1970, 356, pl. 1(4a, 4b, 5a, 5b);
Lacourt 1968
: 72
, pl. 15(j);
Rao 1973
: 529
; 1976, 339, fig. 4H;
Rao
et al
. 1985
: 261
;
Smith & Wood 1995
: 362
, fig. 1;
Wood & Wood 2000
, 423, fig. 1;
Wood
et al
. 2006
: 14
, figs 20–21, 36.
Plumatella emarginata
var.
javanica
:
Loppens 1908
: 162
.
Material examined.
Mature colony with floatoblasts from Fukuji Dam, village of Higashi.
Description.
Observed colony light brown or reddish brown, its surface encrusted (
Fig. 3
A); almost entirely recumbent, with pronounced keel, ends of branches often rising from substratum (
Fig. 3
B). Tentacle number about 45. Floatoblast elliptical in our specimen, large, about 340–410 (366±38) μm long by 180–236 (215±31) μm wide (n=3) (
Fig. 3
C); length/width ratio about 1.8. Fenestrae of floatoblast large; in SEM view, surface entirely covered with minute, rounded tubercles that are rounded-hexagonal in outline, with 5–8 (average 6) small, irregular pores in the angles (
Fig. 3
D). Ventral fenestra 200–246 (225±23) μm long by 182–211 (194±15) μm wide; dorsal fenestra about 150–180 (174±10) μm long by 130–160 (139±16) μm wide (n=3). Floatoblast valves asymmetrical in lateral and transverse views, the ventral valve strongly convex and dorsal valve almost flat. Suture line of both valves with a simple ridge. Sessoblast also with tuberculate surface (
Wood
et al
. 2006
).
Distribution.
Distributed mainly in tropical regions (
Bushnell 1973
), including Southeast Asia (
Lacourt 1968
;
Wood
et al
. 2006
), central Africa (
Wiebach 1964
), and the Amazon River in South
America
(
Wiebach 1967
,
1970
). This is the first record of
P. javanica
from
Japan
, where it occurs on Okinawa and the Pacific side of southern Honshu (Ibaraki, Tokyo, and
Kochi
Prefectures).
Remarks.
Kraepelin (1906)
originally described
P. javanica
from
Indonesia
. Previous authors have considered statoblast morphology, colony color and colony form as important diagnostic characters of this species.
Annandale (1911)
reported a transparent, glassy ectocyst and a strong raphe, and
Smith & Wood (1995)
also indicated these characters. However, as with other plumatellid species, these characters vary ecophenotypically; for example,
Annandale (1911)
specifically mentioned variation in the form of colonies attached to dead wood. Like
P. javanica
,
P. vorstmani
(see below) also has a transparent ectocyst, but the tentacle number in
P. j a v a n i c a
is almost twice that in
P. vorstmani
, and statoblast size and morphology are different.