<document id="B84D0ADA7B56418A8665CDFF4366DDE7" ID-DOI="10.5281/zenodo.5392165" ID-ISSN="1638-9387" ID-Zenodo-Dep="5392165" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630316154645" checkinUser="marcus" docAuthor="Montreuil, Olivier" docDate="2005" docId="03DD87EDFFD4FFAD1944FB8555D9FE29" docLanguage="fr" docName="Zoosystema.2005.27.1.123-135.pdf" docOrigin="Zoosystema 27 (1)" docStyle="DocumentStyle:64A3174DE33E974D4EB6573A0A793B12.4:Zoosystema.2000-2008.journal_article.type2" docStyleId="64A3174DE33E974D4EB6573A0A793B12" docStyleName="Zoosystema.2000-2008.journal_article.type2" docStyleVersion="4" docTitle="Helictopleurus steineri Paulian &amp; Cambefort 1991" docType="treatment" docVersion="4" lastPageNumber="132" masterDocId="FFE4FF95FFDCFFA41A1FFFB7514AFF80" masterDocTitle="Nouveaux Helictopleurus d’Orbigny, 1915 de Madagascar et révision du « groupe semivirens » sensu Lebis, 1960 (Insecta, Coleoptera, Scarabaeidae, Oniticellini)" masterLastPageNumber="135" masterPageNumber="123" pageNumber="131" updateTime="1699223719786" updateUser="plazi" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="3026944F11AA5F37D57D79A94FAF3976">Nouveaux Helictopleurus d’Orbigny, 1915 de Madagascar et révision du « groupe semivirens » sensu Lebis, 1960 (Insecta, Coleoptera, Scarabaeidae, Oniticellini)</mods:title>
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<mods:namePart id="6A8DF8D6275D3BDA4BD7D7E936695678">Montreuil, Olivier</mods:namePart>
<mods:affiliation id="0F497FE54BB67B1915251F61BF5FA1BA">Département Systématique et Évolution, UMR 5202, Muséum national d’Histoire naturelle, Entomologie, case postale 50, 45 rue Buffon, F- 75231 Paris cedex 05 (France) o. montr. @ mnhn. fr ..</mods:affiliation>
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<mods:date id="74777294476F8CAC6A283E95F4B00E21">2005</mods:date>
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<taxonomicName id="4C744D78FFD4FFAC1944FB855527FBEB" ID-CoL="6LKF4" authority="Paulian &amp; Cambefort, 1991" authorityName="Paulian &amp; Cambefort" authorityYear="1991" class="Insecta" family="Scarabaeidae" genus="Helictopleurus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="131" phylum="Arthropoda" rank="species" species="steineri">
<emphasis id="B900EAE9FFD4FFAC1944FB855504FBCB" bold="true" box="[859,1102,1074,1099]" italics="true" pageId="8" pageNumber="131">Helictopleurus steineri</emphasis>
<bibRefCitation id="EFE54B0AFFD4FFAC1923FBE55527FBEB" author="PAULIAN R. &amp; CAMBEFORT Y." box="[828,1133,1106,1131]" pageId="8" pageNumber="131" pagination="113 - 118" refId="ref7470" refString="PAULIAN R. &amp; CAMBEFORT Y. 1991. - Remarques sur le genre Helictopleurus et description de trois nouvelles especes (Coleoptera, Scarabaeidae). Revue francaise d'Entomologie (N. S.) 13: 113 - 118." type="journal article" year="1991">Paulian &amp; Cambefort, 1991</bibRefCitation>
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<treatmentCitation id="0AD510EAFFD4FFAC18D5FBCB53B9FB2D" author="PAULIAN R. &amp; CAMBEFORT Y." page="116" pageId="8" pageNumber="131" year="1991">
<taxonomicName id="4C744D78FFD4FFAC18D5FBCB53B9FB2D" ID-CoL="6LKF4" authority="Paulian &amp; Cambefort, 1991: 116" authorityName="Paulian &amp; Cambefort" authorityPageNumber="116" authorityYear="1991" class="Insecta" family="Scarabaeidae" genus="Helictopleurus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="131" phylum="Arthropoda" rank="species" species="steineri">
<emphasis id="B900EAE9FFD4FFAC18D5FBCB52EFFB13" box="[714,933,1148,1171]" italics="true" pageId="8" pageNumber="131">Helictopleurus steineri</emphasis>
<bibRefCitation id="EFE54B0AFFD4FFAC19B1FBCB53B9FB2D" author="PAULIAN R. &amp; CAMBEFORT Y." pageId="8" pageNumber="131" pagination="113 - 118" refId="ref7470" refString="PAULIAN R. &amp; CAMBEFORT Y. 1991. - Remarques sur le genre Helictopleurus et description de trois nouvelles especes (Coleoptera, Scarabaeidae). Revue francaise d'Entomologie (N. S.) 13: 113 - 118." type="journal article" year="1991">Paulian &amp; Cambefort, 1991: 116</bibRefCitation>
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.
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<paragraph id="8BCB36FBFFD4FFAC18D5FB095582FAA1" blockId="8.[714,1247,1074,1313]" pageId="8" pageNumber="131">
MATÉRIEL TYPE. — 
<typeStatus id="54CF8859FFD4FFAC19D9FB09557AFB55" box="[966,1072,1214,1237]" pageId="8" pageNumber="131" type="holotype">Holotype</typeStatus>
: 
<collectingCountry id="F363766BFFD4FFAC1E4CFB09559DFB55" box="[1107,1239,1214,1237]" name="Madagascar" pageId="8" pageNumber="131">Madagascar</collectingCountry>
, Ranomafana, 
<collectingRegion id="49B0F819FFD4FFAC1944FB6F5528FB6F" box="[859,1122,1240,1263]" country="Madagascar" name="Fianarantsoa" pageId="8" pageNumber="131">Province de Fianarantsoa</collectingRegion>
, (Smithsonian Institution, Washington) (non examiné). 
<typeStatus id="54CF8859FFD4FFAC18D5FABD526DFAA1" box="[714,807,1290,1313]" pageId="8" pageNumber="131" type="paratype">Paratypes</typeStatus>
: mêmes données, 
<specimenCount id="9D72FD72FFD4FFAC19FEFABD551DFAA1" box="[993,1111,1290,1313]" pageId="8" pageNumber="131" type="generic">3 spécimens</specimenCount>
(MNHN).
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<paragraph id="8BCB36FBFFD4FFAC18D5FAE55211FAEA" blockId="8.[714,968,1362,1451]" box="[714,859,1362,1387]" pageId="8" pageNumber="131">DESCRIPTION</paragraph>
<paragraph id="8BCB36FBFFD4FFAC18D5FAC55282FA0B" blockId="8.[714,968,1362,1451]" box="[714,968,1394,1419]" pageId="8" pageNumber="131">
Édéage: voir 
<figureCitation id="134F2A7EFFD4FFAC1942FAC5528EFA0B" box="[861,964,1394,1419]" captionStart="FIG" captionStartId="4.[151,162,1468,1485]" captionTargetBox="[226,1166,246,1428]" captionTargetId="figure-202@4.[219,1181,242,1450]" captionTargetPageId="4" captionText="FIG. 2. — Édéage des Helictopleurus du « groupe semivirens », paramère gauche, vue dorsale et paramère droit; A, H. dorbignyi n. sp.; B, H. tamatavensis n. sp.; C, H. cambeforti n. sp.; D, H. pseudoniger n. sp.; E, H. niger d’Orbigny, 1915; F, H. seminiger d’Orbigny, 1915; G, H. aeneoniger Lebis, 1960; H, H. semivirens d’Orbigny, 1915; I, H. steineri Paulian &amp; Cambefort, 1991. Échelle: A-H, 1 mm; I, 0,5 mm." figureDoi="http://doi.org/10.5281/zenodo.5392169" httpUri="https://zenodo.org/record/5392169/files/figure.png" pageId="8" pageNumber="131">Figure 2I</figureCitation>
.
</paragraph>
<paragraph id="8BCB36FBFFD4FFAC18D5FA2552E6FA2B" blockId="8.[714,968,1362,1451]" box="[714,940,1426,1451]" pageId="8" pageNumber="131">
Tête: voir 
<figureCitation id="134F2A7EFFD4FFAC195EFA2552E2FA2B" box="[833,936,1426,1451]" captionStart="FIG" captionStartId="6.[151,162,1681,1698]" captionTargetBox="[321,1089,240,1642]" captionTargetId="figure-24@6.[314,1118,213,1672]" captionTargetPageId="6" captionText="FIG. 3. — Tête des Helictopleurus du « groupe semivirens », mâle, vue frontale; A, H. dorbignyi n. sp.; B, H. tamatavensis n. sp.; C, H. cambeforti n. sp.; D, H. pseudoniger n. sp.; E, H. niger d’Orbigny, 1915; F, H. seminiger d’Orbigny, 1915; G, H. aeneoniger" figureDoi="http://doi.org/10.5281/zenodo.5392171" httpUri="https://zenodo.org/record/5392171/files/figure.png" pageId="8" pageNumber="131">Figure 3I</figureCitation>
.
</paragraph>
<paragraph id="8BCB36FBFFD4FFAC18D5FA65558EF98B" blockId="8.[714,1247,1490,1547]" pageId="8" pageNumber="131">
La répartition des neuf espèces du « groupe 
<taxonomicName id="4C744D78FFD4FFAC1EB6FA65524CF98B" authorityName="d'Orbigny" authorityYear="1915" class="Insecta" family="Scarabaeidae" genus="Helictopleurus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="131" phylum="Arthropoda" rank="species" species="semivirens">
<emphasis id="B900EAE9FFD4FFAC1EB6FA65524CF98B" italics="true" pageId="8" pageNumber="131">semivirens</emphasis>
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» est présentée sur la carte de la 
<figureCitation id="134F2A7EFFD4FFAC1E7CFA45558AF98B" box="[1123,1216,1522,1547]" captionStart="FIG" captionStartId="8.[714,725,870,887]" captionTargetBox="[837,1140,247,830]" captionTargetId="figure-789@8.[819,1140,233,832]" captionTargetPageId="8" captionText="FIG. 4. — Répartition des Helictopleurus du « groupe semivirens »::, H. aeneoniger Lebis, 1960; Ë, H. cambeforti n. sp.; ✫, H. tamatavensis n. sp.; O, H. pseudoniger n. sp.; ▲, H. dorbignyi n. sp.; ●, H. semivirens d’Orbigny, 1915; Î, H. steineri Paulian &amp; Cambefort, 1991; ■, H. niger d’Orbigny, 1915; +, H. seminiger d’Orbigny, 1915." figureDoi="http://doi.org/10.5281/zenodo.5392173" httpUri="https://zenodo.org/record/5392173/files/figure.png" pageId="8" pageNumber="131">Figure 4</figureCitation>
.
</paragraph>
<paragraph id="8BCB36FBFFD5FFAD1AB9FFC35043FF05" blockId="9.[166,265,116,133]" box="[166,265,116,133]" pageId="9" pageNumber="132">Montreuil O.</paragraph>
<paragraph id="8BCB36FBFFD5FFAD1B2FFF45550CFE8B" blockId="9.[151,1246,242,363]" box="[304,1094,242,267]" pageId="9" pageNumber="132">
CLÉ DE DÉTERMINATION DES 
<taxonomicName id="4C744D78FFD5FFAD1872FF455200FE8A" authority="DU" authorityName="DU" box="[621,842,242,267]" class="Insecta" family="Scarabaeidae" genus="Helictopleurus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="132" phylum="Arthropoda" rank="genus">
<emphasis id="B900EAE9FFD5FFAD1872FF455269FE8A" box="[621,803,242,267]" italics="true" pageId="9" pageNumber="132">HELICTOPLEURUS</emphasis>
DU
</taxonomicName>
« GROUPE 
<taxonomicName id="4C744D78FFD5FFAD19A0FF40557EFE8A" authorityName="d'Orbigny" authorityYear="1915" box="[959,1076,247,266]" class="Insecta" family="Scarabaeidae" genus="Helictopleurus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="132" phylum="Arthropoda" rank="species" species="semivirens">
<emphasis id="B900EAE9FFD5FFAD19A0FF40557EFE8A" box="[959,1076,247,266]" italics="true" pageId="9" pageNumber="132">SEMIVIRENS</emphasis>
</taxonomicName>
»
</paragraph>
<paragraph id="8BCB36FBFFD5FFAD1A88FE85500AFEEB" blockId="9.[151,1246,242,363]" pageId="9" pageNumber="132">(Les femelles étant inconnues pour la plus grande partie des espèces de ce groupe, cette clé est donnée pour les mâles).</paragraph>
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1. Espèce entièrement noire, sans reflet métallique........................................ 
<taxonomicName id="4C744D78FFD5FFAD1E39FE2755D9FE29" ID-CoL="6LKF4" authorityName="Paulian &amp; Cambefort" authorityYear="1991" box="[1062,1171,400,425]" class="Insecta" family="Scarabaeidae" genus="Helictopleurus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="132" phylum="Arthropoda" rank="species" species="steineri">
<emphasis id="B900EAE9FFD5FFAD1E39FE2755D9FE29" box="[1062,1171,400,425]" italics="true" pageId="9" pageNumber="132">H. steineri</emphasis>
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