<document id="AF25FF923196874AFF0A69E34C2C887F" ID-CLB-Dataset="7890" ID-DOI="10.11646/zootaxa.4871.1.1" ID-GBIF-Dataset="344f8a86-21a1-428e-ae4f-01ea6082254a" ID-ISSN="1175-5326" ID-Zenodo-Dep="4423340" ID-ZooBank="C21A719F-9A6B-4227-8386-1AFA22620614" IM.bibliography_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1610022441531" checkinUser="plazi" docAuthor="Guglielmone, Alberto A., Petney, Trevor N. &amp; Robbins, Richard G." docDate="2020" docId="C04787D4FF4CFF64FF07F8396169C97A" docLanguage="en" docName="zootaxa.4871.1.1.pdf" docOrigin="Zootaxa 4871 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Haemaphysalis howletti Warburton 1913" docType="treatment" docVersion="9" lastPageNumber="167" masterDocId="3C7EFFACFFE9FFC2FF90FFA76543CB47" masterDocTitle="Ixodidae (Acari: Ixodoidea): descriptions and redescriptions of all known species from 1758 to December 31, 2019" masterLastPageNumber="322" masterPageNumber="1" pageNumber="166" updateTime="1698923194747" updateUser="ExternalLinkService">
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<mods:title id="7DFDF8955A445A8F820E0C749FAE92E1">Ixodidae (Acari: Ixodoidea): descriptions and redescriptions of all known species from 1758 to December 31, 2019</mods:title>
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<mods:namePart id="6D9B9E8D84ADECD89A0A43DBF25FD6EA">Guglielmone, Alberto A.</mods:namePart>
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<mods:namePart id="50D91A92E90FE265FD7CD26CAC9FF6F7">Petney, Trevor N.</mods:namePart>
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<mods:namePart id="876289C3695DCDDBABE9749B87D4FDF2">Robbins, Richard G.</mods:namePart>
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<treatment id="C04787D4FF4CFF64FF07F8396169C97A" ID-DOI="http://doi.org/10.5281/zenodo.4583462" ID-GBIF-Taxon="179990253" ID-Zenodo-Dep="4583462" LSID="urn:lsid:plazi:treatment:C04787D4FF4CFF64FF07F8396169C97A" httpUri="http://treatment.plazi.org/id/C04787D4FF4CFF64FF07F8396169C97A" lastPageId="166" lastPageNumber="167" pageId="165" pageNumber="166">
<subSubSection id="00F46549FF4CFF67FF07F83967EACCFF" box="[151,681,1950,1977]" pageId="165" pageNumber="166" type="nomenclature">
<paragraph id="485136C2FF4CFF67FF07F83967EACCFF" blockId="165.[151,1436,1950,2013]" box="[151,681,1950,1977]" pageId="165" pageNumber="166">
<emphasis id="7A9AEAD0FF4CFF67FF07F83965F9CCFF" bold="true" box="[151,186,1950,1976]" pageId="165" pageNumber="166">63.</emphasis>
<taxonomicName id="8FEE4D41FF4CFF67FF57F83967E6CCFF" ID-CoL="3J7W7" authority="Warburton, 1913" authorityName="Warburton" authorityYear="1913" box="[199,677,1950,1977]" class="Arachnida" family="Ixodidae" genus="Haemaphysalis" kingdom="Animalia" order="Ixodida" pageId="165" pageNumber="166" phylum="Arthropoda" rank="species" species="howletti">
<emphasis id="7A9AEAD0FF4CFF67FF57F839649FCCFE" bold="true" box="[199,476,1950,1977]" italics="true" pageId="165" pageNumber="166">Haemaphysalis howletti</emphasis>
<bibRefCitation id="2C7F4B33FF4CFF67FE74F83967E6CCFF" author="Warburton, C." box="[484,677,1950,1976]" pageId="165" pageNumber="166" pagination="121 - 130" refId="ref254599" refString="Warburton, C. (1913) On four new species and two varieties of the ixodid genus Haemaphysalis. Parasitology, 6, 121 - 130. https: // doi. org / 10.1017 / S 0031182000002973" type="journal article" year="1913">Warburton, 1913</bibRefCitation>
</taxonomicName>
.
</paragraph>
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<subSubSection id="00F46549FF4CFF67FD3FF83960DFCC9B" pageId="165" pageNumber="166" type="discussion">
<paragraph id="485136C2FF4CFF67FD3FF83960DFCC9B" blockId="165.[151,1436,1950,2013]" pageId="165" pageNumber="166">
An 
<collectingRegion id="8A2AF820FF4CFF67FD49F8396676CCFF" box="[729,821,1950,1976]" country="Saudi Arabia" name="Ash Sharqiyah" pageId="165" pageNumber="166">Oriental</collectingRegion>
species, all of whose parasitic stages have been found on 
<taxonomicName id="8FEE4D41FF4CFF67FF2DF8656464CC9B" box="[189,295,1986,2012]" class="Mammalia" kingdom="Animalia" order="Rodentia" pageId="165" pageNumber="166" phylum="Chordata" rank="order">Rodentia</taxonomicName>
: 
<taxonomicName id="8FEE4D41FF4CFF67FEA3F86564D5CC9B" box="[307,406,1986,2012]" class="Mammalia" family="Muridae" kingdom="Animalia" order="Rodentia" pageId="165" pageNumber="166" phylum="Chordata" rank="family">Muridae</taxonomicName>
; adults have been recovered from 
<taxonomicName id="8FEE4D41FF4CFF67FCB6F86566F9CC9B" box="[806,954,1986,2012]" class="Mammalia" kingdom="Animalia" order="Lagomorpha" pageId="165" pageNumber="166" phylum="Chordata" rank="order">Lagomorpha</taxonomicName>
: 
<taxonomicName id="8FEE4D41FF4CFF67FC56F8656179CC9B" box="[966,1082,1986,2012]" class="Mammalia" family="Leporidae" kingdom="Animalia" order="Lagomorpha" pageId="165" pageNumber="166" phylum="Chordata" rank="family">Leporidae</taxonomicName>
, and Persissodactyla: 
<taxonomicName id="8FEE4D41FF4CFF67FAA9F86560DBCC9B" authorityName="Gray" authorityYear="1821" box="[1337,1432,1986,2012]" class="Mammalia" family="Equidae" kingdom="Animalia" order="Perissodactyla" pageId="165" pageNumber="166" phylum="Chordata" rank="family">Equidae</taxonomicName>
;
</paragraph>
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<subSubSection id="00F46549FF4FFF64FF07FF306169C97A" pageId="166" pageNumber="167" type="description">
<paragraph id="485136C2FF4FFF64FF07FF3066D8CB92" blockId="166.[151,1436,151,573]" pageId="166" pageNumber="167">
nymphs have been collected from 
<taxonomicName id="8FEE4D41FF4FFF64FD89FF3067F5CBF6" box="[537,694,151,177]" class="Aves" kingdom="Animalia" order="Cuculiformes" pageId="166" pageNumber="167" phylum="Chordata" rank="order">Cuculiformes</taxonomicName>
: 
<taxonomicName id="8FEE4D41FF4FFF64FD50FF306676CBF6" box="[704,821,151,177]" class="Aves" family="Cuculidae" kingdom="Animalia" order="Cuculiformes" pageId="166" pageNumber="167" phylum="Chordata" rank="family">Cuculidae</taxonomicName>
; larvae have been found on 
<taxonomicName id="8FEE4D41FF4FFF64FBE0FF306199CBF6" box="[1136,1242,151,177]" class="Mammalia" kingdom="Animalia" order="Rodentia" pageId="166" pageNumber="167" phylum="Chordata" rank="order">Rodentia</taxonomicName>
: 
<taxonomicName id="8FEE4D41FF4FFF64FB74FF306012CBF6" box="[1252,1361,151,177]" class="Mammalia" family="Sciuridae" kingdom="Animalia" order="Rodentia" pageId="166" pageNumber="167" phylum="Chordata" rank="family">Sciuridae</taxonomicName>
. There are no records of 
<taxonomicName id="8FEE4D41FF4FFF64FECBFF1C672ACB92" authorityName="Warburton" authorityYear="1913" box="[347,617,187,213]" class="Arachnida" family="Ixodidae" genus="Haemaphysalis" kingdom="Animalia" order="Ixodida" pageId="166" pageNumber="167" phylum="Arthropoda" rank="species" species="howletti">
<emphasis id="7A9AEAD0FF4FFF64FECBFF1C672ACB92" box="[347,617,187,213]" italics="true" pageId="166" pageNumber="167">Haemaphysalis howletti</emphasis>
</taxonomicName>
causing human parasitism.
</paragraph>
<paragraph id="485136C2FF4FFF64FF07FF7864C9CBBE" blockId="166.[151,1436,151,573]" box="[151,394,223,249]" pageId="166" pageNumber="167">
<emphasis id="7A9AEAD0FF4FFF64FF07FF7865F8CBBE" bold="true" box="[151,187,223,249]" pageId="166" pageNumber="167">M:</emphasis>
<bibRefCitation id="2C7F4B33FF4FFF64FF52FF7864C9CBBE" author="Warburton, C." box="[194,394,223,249]" pageId="166" pageNumber="167" pagination="121 - 130" refId="ref254599" refString="Warburton, C. (1913) On four new species and two varieties of the ixodid genus Haemaphysalis. Parasitology, 6, 121 - 130. https: // doi. org / 10.1017 / S 0031182000002973" type="journal article" year="1913">Warburton (1913)</bibRefCitation>
</paragraph>
<paragraph id="485136C2FF4FFF64FF07FEA464C2CA5A" blockId="166.[151,1436,151,573]" box="[151,385,259,285]" pageId="166" pageNumber="167">
<emphasis id="7A9AEAD0FF4FFF64FF07FEA465F1CA5A" bold="true" box="[151,178,259,285]" pageId="166" pageNumber="167">F:</emphasis>
<bibRefCitation id="2C7F4B33FF4FFF64FF29FEA464C2CA5A" author="Warburton, C." box="[185,385,259,285]" pageId="166" pageNumber="167" pagination="121 - 130" refId="ref254599" refString="Warburton, C. (1913) On four new species and two varieties of the ixodid genus Haemaphysalis. Parasitology, 6, 121 - 130. https: // doi. org / 10.1017 / S 0031182000002973" type="journal article" year="1913">Warburton (1913)</bibRefCitation>
</paragraph>
<paragraph id="485136C2FF4FFF64FF07FE806432CA05" blockId="166.[151,1436,151,573]" box="[151,369,295,322]" pageId="166" pageNumber="167">
<emphasis id="7A9AEAD0FF4FFF64FF07FE8065F6CA06" bold="true" box="[151,181,295,321]" pageId="166" pageNumber="167">N:</emphasis>
<bibRefCitation id="2C7F4B33FF4FFF64FF2CFE806432CA05" author="Dhanda, V." box="[188,369,295,322]" pageId="166" pageNumber="167" pagination="459 - 465" refId="ref194594" refString="Dhanda, V. (1964 a) Description of immature stages of Haemaphysalis howletti (Ixodoidea: Ixodidae), and redescription of adults. Journal of Parasitology, 50, 459 - 465. https: // doi. org / 10.2307 / 3275856" type="journal article" year="1964">Dhanda (1964a)</bibRefCitation>
</paragraph>
<paragraph id="485136C2FF4FFF64FF07FEEC642CCA21" blockId="166.[151,1436,151,573]" box="[151,367,331,358]" pageId="166" pageNumber="167">
<emphasis id="7A9AEAD0FF4FFF64FF07FEEC65F0CA22" bold="true" box="[151,179,331,357]" pageId="166" pageNumber="167">L:</emphasis>
<bibRefCitation id="2C7F4B33FF4FFF64FF2AFEEC642CCA21" author="Dhanda, V." box="[186,367,331,358]" pageId="166" pageNumber="167" pagination="459 - 465" refId="ref194594" refString="Dhanda, V. (1964 a) Description of immature stages of Haemaphysalis howletti (Ixodoidea: Ixodidae), and redescription of adults. Journal of Parasitology, 50, 459 - 465. https: // doi. org / 10.2307 / 3275856" type="journal article" year="1964">Dhanda (1964a)</bibRefCitation>
</paragraph>
<paragraph id="485136C2FF4FFF64FF07FEC9640BCACF" blockId="166.[151,1436,151,573]" box="[151,328,366,392]" pageId="166" pageNumber="167">
<emphasis id="7A9AEAD0FF4FFF64FF07FEC9640BCACF" bold="true" box="[151,328,366,392]" pageId="166" pageNumber="167">Redescriptions</emphasis>
</paragraph>
<paragraph id="485136C2FF4FFF64FF07FE34668DCAEA" blockId="166.[151,1436,151,573]" box="[151,974,403,430]" pageId="166" pageNumber="167">
<emphasis id="7A9AEAD0FF4FFF64FF07FE3465F8CAEA" bold="true" box="[151,187,403,429]" pageId="166" pageNumber="167">M:</emphasis>
<bibRefCitation id="2C7F4B33FF4FFF64FF52FE3464D2CAEA" author="Toumanoff, C." box="[194,401,403,429]" pageId="166" pageNumber="167" refId="ref251642" refString="Toumanoff, C. (1944) Les tiques (Ixodoidea) de l'Indochine. Institut Pasteur de l'Indochine, S. I. L. I., Saigon, 220 pp." type="book" year="1944">Toumanoff (1944)</bibRefCitation>
, 
<bibRefCitation id="2C7F4B33FF4FFF64FE0DFE346716CAE9" author="Dhanda, V." box="[413,597,403,430]" pageId="166" pageNumber="167" pagination="459 - 465" refId="ref194594" refString="Dhanda, V. (1964 a) Description of immature stages of Haemaphysalis howletti (Ixodoidea: Ixodidae), and redescription of adults. Journal of Parasitology, 50, 459 - 465. https: // doi. org / 10.2307 / 3275856" type="journal article" year="1964">Dhanda (1964a)</bibRefCitation>
, 
<bibRefCitation id="2C7F4B33FF4FFF64FDF0FE346689CAEA" author="Geevarghese, G. &amp; Mishra, A. C." box="[608,970,403,429]" pageId="166" pageNumber="167" refId="ref202939" refString="Geevarghese, G. &amp; Mishra, A. C. (2011) Haemaphysalis ticks of India. Elsevier, Amsterdam, 260 pp." type="book" year="2011">Geevarghese and Mishra (2011)</bibRefCitation>
.
</paragraph>
<paragraph id="485136C2FF4FFF64FF07FE106686CA96" blockId="166.[151,1436,151,573]" box="[151,965,439,466]" pageId="166" pageNumber="167">
<emphasis id="7A9AEAD0FF4FFF64FF07FE1065F1CA96" bold="true" box="[151,178,439,465]" pageId="166" pageNumber="167">F:</emphasis>
<bibRefCitation id="2C7F4B33FF4FFF64FF29FE1064CBCA96" author="Toumanoff, C." box="[185,392,439,465]" pageId="166" pageNumber="167" refId="ref251642" refString="Toumanoff, C. (1944) Les tiques (Ixodoidea) de l'Indochine. Institut Pasteur de l'Indochine, S. I. L. I., Saigon, 220 pp." type="book" year="1944">Toumanoff (1944)</bibRefCitation>
, 
<bibRefCitation id="2C7F4B33FF4FFF64FE04FE106708CA95" author="Dhanda, V." box="[404,587,439,466]" pageId="166" pageNumber="167" pagination="459 - 465" refId="ref194594" refString="Dhanda, V. (1964 a) Description of immature stages of Haemaphysalis howletti (Ixodoidea: Ixodidae), and redescription of adults. Journal of Parasitology, 50, 459 - 465. https: // doi. org / 10.2307 / 3275856" type="journal article" year="1964">Dhanda (1964a)</bibRefCitation>
, 
<bibRefCitation id="2C7F4B33FF4FFF64FDC7FE106683CA96" author="Geevarghese, G. &amp; Mishra, A. C." box="[599,960,439,465]" pageId="166" pageNumber="167" refId="ref202939" refString="Geevarghese, G. &amp; Mishra, A. C. (2011) Haemaphysalis ticks of India. Elsevier, Amsterdam, 260 pp." type="book" year="2011">Geevarghese and Mishra (2011)</bibRefCitation>
.
</paragraph>
<paragraph id="485136C2FF4FFF64FF07FE7C6761CAB2" blockId="166.[151,1436,151,573]" box="[151,546,475,501]" pageId="166" pageNumber="167">
<emphasis id="7A9AEAD0FF4FFF64FF07FE7C65F6CAB2" bold="true" box="[151,181,475,501]" pageId="166" pageNumber="167">N:</emphasis>
<bibRefCitation id="2C7F4B33FF4FFF64FF2CFE7C6761CAB2" author="Geevarghese, G. &amp; Mishra, A. C." box="[188,546,475,501]" pageId="166" pageNumber="167" refId="ref202939" refString="Geevarghese, G. &amp; Mishra, A. C. (2011) Haemaphysalis ticks of India. Elsevier, Amsterdam, 260 pp." type="book" year="2011">Geevarghese and Mishra (2011)</bibRefCitation>
</paragraph>
<paragraph id="485136C2FF4FFF64FF07FE586762C95E" blockId="166.[151,1436,151,573]" box="[151,545,511,537]" pageId="166" pageNumber="167">
<emphasis id="7A9AEAD0FF4FFF64FF07FE5865F0C95E" bold="true" box="[151,179,511,537]" pageId="166" pageNumber="167">L:</emphasis>
<bibRefCitation id="2C7F4B33FF4FFF64FF2AFE586762C95E" author="Geevarghese, G. &amp; Mishra, A. C." box="[186,545,511,537]" pageId="166" pageNumber="167" refId="ref202939" refString="Geevarghese, G. &amp; Mishra, A. C. (2011) Haemaphysalis ticks of India. Elsevier, Amsterdam, 260 pp." type="book" year="2011">Geevarghese and Mishra (2011)</bibRefCitation>
</paragraph>
<paragraph id="485136C2FF4FFF64FF07FD846169C97A" blockId="166.[151,1436,151,573]" box="[151,1066,547,573]" pageId="166" pageNumber="167">
<emphasis id="7A9AEAD0FF4FFF64FF07FD84659BC97A" bold="true" box="[151,216,547,573]" pageId="166" pageNumber="167">Note:</emphasis>
see 
<taxonomicName id="8FEE4D41FF4FFF64FE9AFD846754C97A" box="[266,535,547,573]" class="Arachnida" family="Ixodidae" genus="Haemaphysalis" kingdom="Animalia" order="Ixodida" pageId="166" pageNumber="167" phylum="Arthropoda" rank="species" species="doenitzi">
<emphasis id="7A9AEAD0FF4FFF64FE9AFD846754C97A" box="[266,535,547,573]" italics="true" pageId="166" pageNumber="167">Haemaphysalis doenitzi</emphasis>
</taxonomicName>
for its confusion with 
<emphasis id="7A9AEAD0FF4FFF64FC85FD846169C97A" box="[789,1066,547,573]" italics="true" pageId="166" pageNumber="167">
<taxonomicName id="8FEE4D41FF4FFF64FC85FD846165C97A" authorityName="Warburton" authorityYear="1913" box="[789,1062,547,573]" class="Arachnida" family="Ixodidae" genus="Haemaphysalis" kingdom="Animalia" order="Ixodida" pageId="166" pageNumber="167" phylum="Arthropoda" rank="species" species="howletti">Haemaphysalis howletti</taxonomicName>
.
</emphasis>
</paragraph>
</subSubSection>
</treatment>
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