Lost and found: The Eocene family Pyramimitridae (Neogastropoda) discovered in the Recent fauna of the Indo-Pacific
Author
Kantor, Yuri
Author
Lozouet, Pierre
Author
Puillandre, Nicolas
Author
Bouchet, Philippe
text
Zootaxa
2014
3754
3
239
276
journal article
46667
10.11646/zootaxa.3754.3.2
fe375541-8083-43f8-9cf8-b8d75af79599
1175-5326
251551
9E645014-5464-4E7C-8D4A-0B3B52A5AA53
Vaughanites
Woodring, 1928
Vaughanites leptus
Woodring, 1928
Miocene
,
Jamaica
.
Vaughanites
?
superstes
new species
Philippines
and
Papua New Guinea
,
318–
389 m
.
Until the present paper, the family would have been thought to be restricted to Eocene-Miocene fossils. However, based on the number of known fossil (8) and Recent (11) species, it is clear that we are not merely dealing with the survivors of a once diversified family. Indeed, the Recent fauna is richer than the fossil ones and it had merely been overlooked. This can be explained by their occurrence in relatively deep water, their scarcity, and their rather non-descript "small turrid" appearance which makes them look like one of thousands of undescribed turrid species. Additional new Recent and fossil species may already be lying buried in the collections of deepwater unsorted turrids in museums.
Among the fossils, the extinct genera
Pyramimitra
and
Endiatoma
originate from shallow water deposits, and the Lower Eocene
Endiatoma
represents the oldest occurrence of the family. Conversely, the two fossil
Hortia
from the Upper Eocene and the Upper Oligocene come from offshore, upper bathyal deposits (
Lozouet 2004
).
All fossil
Pyramimitridae
have a multispiral protoconch of 2.5–3.5 whorls, indicating planktotrophic larval development. Sculpture of axial ribs, appearing on the second or third whorl, is reminiscent of that observed in modern (and fossil) planktotrophic species of
Fasciolariidae
. By contrast, all but two Recent species possess a large, bulbous, paucispiral protoconch, indicating non-planktotrophic development that may be correlated with narrow geographical ranges. The exceptions are
Teremitra efatensis
and
Vaughanites
?
superstes
that have a multispiral protoconch and broad distributions extending from the
Philippines
to
Fiji
, and the
Philippines
to
Papua New Guinea
, respectively.