The genus Gromphas Dejean, 1836 (Coleoptera, Scarabaeinae): nomenclature, distribution, and conservation, including a contribution to the debate on electronic publications in zoology Author Cupello, Mario Laboratório de Sistemática e Bioecologia de Coleoptera, Departamento de Zoologia, Universidade Federal do Paraná, Caixa Postal 19020, 81531 - 980, Curitiba, Paraná (Brazil); and Department of Entomology, Texas A & M University, College Station, TX 77843 (United States) mcupello @ hotmail. com mcupello@hotmail.com text Zoosystema 2024 2024-02-09 46 2 23 59 https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/zoosystema2024v46a2.pdf journal article 10.5252/zoosystema2024v46a2 1638-9387 4B49C1D9-1196-4942-969F-2E923B1FC12C Gromphas lemoinei Waterhouse, 1891 Gromphas lemoinei Waterhouse, 1891: 60 [ 1st July 1891 ]. NAME-BEARING TYPE . — Holotype by monotypy (unknown sex), NHM. TYPE LOCALITY . — Venezuela : La Guaira: La Guaira. ETYMOLOGY . — The masculine singular genitive of the proper noun Lemoine, a French surname meaning ‘the monk’ (‘Le Moine’). Waterhouse (1891) did not explain after whom this eponym was formed, but I suspect it may have been the French entomologist Victor Lemoine (b.?-1897), the only Lemoine who I found associated with entomology in the late 19th century. Alas, apart from the fact that he was based in Paris and was a pioneer in the application of radiography in the study of arthropods ( Lemoine 1897 ), almost nothing is known about Victor Lemoine ( Anonymous 1897 ; Constantin 2012 ; Groll 2016 ). DISTRIBUTION . — Open savannas, pastures and river sandbanks in the Colombian and Venezuelan Llanos and associated areas.