Trapdoor Spiders of the Genus Misgolas (Mygalomorphae: Idiopidae) from Eastern New South Wales, With Notes on Genetic Variation Author Wishart, Graham Author Rowell, David M. text Records of the Australian Museum 2008 2008-06-25 60 1 45 86 http://dx.doi.org/10.3853/j.0067-1975.60.2008.1495 journal article 10.3853/j.0067-1975.60.2008.1495 2201-4349 5240070 Misgolas helensmithae n.sp. Figs 8A–J , 24A Type material . HOLOTYPE ? , AM KS48654 , Bulga State Forest , Homewoods Road , 2.8 km West of Knodingbul Road ( 31°37'S 152°07'E ), 690 m , 4 Feb.–9 Apr. 1993 , pit fall trap , site 57CR, M. Gray , G. Cassis . PARATYPES ?? : AM KS38642 , AM KS92875 , AM KS92876 , for all paratypes data as for holotype . Diagnosis . In male: medium-large sized dark tan spider, carapace length c. 7.3–8.4; retrodorsal surface of metatarsi IV with spines ( Fig. 8I ); venter entirely pallid ( Fig. 8J ). Tarsi I and II with bilateral rows of small spines adjacent to scopula; metatarsi I and II with bilateral distal clusters of spines ( Fig. 8E–H ). Palpal bulb ( Fig. 8B,C ) with retrolateral embolic flange with c. 20 minute folds; embolus straight from dorsal aspect, apophysis absent. Retrolateral tibial apophysis digitate; palpal tibia as figured ( Fig. 8A ). Female not known. Fig. 8. Misgolas helensmithae n.sp. A–H,J ? , holotype AM KS48654: ( A ), right palp retrolateral. ( B,C ), right bulb: B , dorsal; C , prolateral. ( D ), right cymbium dorsal. ( E,F ), right tarsus/metatarsus I: E , prolateral; F , retrolateral. ( G,H ), right tarsus/metatarsus II: G , prolateral; H , retrolateral. ( J ), venter. ( I ) ? , paratype AM KS38642: right tarsus/metatarsus IV retrolateral. Description Male holotype ( Fig. 8A–J ). Size . Carapace length 8.42, width 6.66. Abdomen length 6.63, width 4.42. Colour . In alcohol carapace, chelicerae and limbs dark tan, v surfaces much paler. Dark brown smudges absent on limbs. Sparse cover of prostrate pallid hairs on carapace, absent on limbs. Abdomen dorsum mottled dark brown bearing c. 5 pallid transverse chevrons. Venter entirely pallid. ( Fig. 8J ). Carapace . Edge fringed with black bristles which encroach onto posterior fifth of post foveal surface. Few small black bristles scattered on caput; line of 6 median bristles on caput arch; 2 bristles between PME, 5 bristles on clypeus; 3 small bristles on weakly chitinous pleuron below clypeus. Fovea width 1.51, gently recurved; posterior wall not pitted. Eyes . Raised on mound. Area immediately adjacent to eyes black excepting between ALE and between PME. Anterior width 1.23, posterior width 1.15, length 0.79, width/length ratio 1.56. Line joining posterior edges of ALE bisects AME. Posterior row straight in front, recurved behind. Chelicerae . Rastellum row of 5(6) spines; few smaller spines receding along anteroental edge of chelicerae. Intercheliceral tumescence distinct, white, not raised, bearing sparse cover of minute brown prostrate setae. Fang groove with 9(18) promarginal teeth and 15(13) smaller retromarginal/intermediate row teeth. Labium . Bulbous, length 0.74, width 1.41. Labio-sternal suture broad, narrowing centrally, divided. Maxillae c. 17(21) anteroental long weak attenuate cuspules. Sternum . Length 4.42, width 3.07. Sigilla all small, round: Anterior pairs submarginal, posterior pair twice diameter from margin. Legs . Tibia I with distal bifid apophysis. Distal process with 1(2) short spine. Proximal process with 3(3) longer spines; 1(1) megaspine rises from d surface of base of proximal process. Palp I II III IV femur 3.99 7.00 6.69 6.14 7.74 patella 1.84 3.25 3.07 2.82 3.68 tibia 3.80 5.34 5.16 3.87 6.94 metatarsus — 6.08 5.53 5.22 7.18 tarsus 1.66 3.50 3.31 3.13 3.81 total 11.29 25.17 23.76 21.18 29.35 Palp ( Fig. 8A,D ). Cymbium with c. 32, many broken away, long, attenuate, anteriorly inclined spines distributed over distal third of d surface. RTA digitate centrally slightly swollen, d and rd surfaces covered with squat pointed spines; proximal half of rv edge of tibial excavation suspending c. 23 similar spines; distal half with a sparse brush of c. 15 longer spines terminating at end of short DTA. Small pointed TEM ( Fig. 8A ) with faint TET adjacent to RTA. Bulb . ( Fig. 8B,C ) Embolic rl flange with c. 21 minute folds, margin distally sharply convexly curved under embolus as if twisted. Embolus straight from d aspect, apophysis absent. Scopula . Complete on tarsi I and II, sparse distal remnants on metatarsi I and II, absent on legs III and IV. Trichobothria . Palp: tarsi 9, tibia pd 5, rd 5. Leg I: tarsi 10, metatarsi 12, tibia pd 5, rd 5. Leg II: tarsi 11, metatarsi 11, tibia pd 6, rd 6. Leg III: tarsi 12, metatarsi 10, tibia pd 6, rd 7. Leg IV: tarsi 11, metatarsi 15, tibia pd 7, rd 7. Leg spination . Leg I ( Fig. 8E,F ): tarsi pv line of 9 adjacent scopula margin, rv line of 17 adjacent scopula margin; metatarsi pv distal cluster of 12, rv cluster of 12 on distal three-quarters; tibia v 0101123, pd 01010; patella pd 1. Leg II ( Fig. 8G,H ): tarsi pv line of 12 adjacent scopula margin, rv line of 14 adjacent scopula margin; metatarsi pv cluster of 9 on distal two-thirds, rv cluster of 14 on distal four-fifths, pd 010; tibia v 011114, pd 011110; patella pd 1. Leg III: tarsi v 25 scattered; metatarsi v 14 scattered, d 002220; tibia v 7 scattered, pl 0110, rl 0110; patella pd 4. Leg IV ( Fig. 8I ): tarsi v 24 scattered; metatarsi v 13 scattered, rd 001010; tibia v 7 scattered. Abdomen . Median band of c. 12 long bristles on dorsum; sparse cover short brown bristles with underlying prostrate setae. Venter covered with small brown bristles. Etymology . The species is named in recognition of arachnologist Dr Helen Smith of the Australian Museum, Sydney. Distribution and natural history . ( Fig. 24A ) This species is known only from the type locality, the Bulga State Forest, c. 30 km ENE of the town of Comboyne, NSW. The burrow is unknown.