Trapdoor Spiders of the Genus Misgolas (Mygalomorphae: Idiopidae) from Eastern New South Wales, With Notes on Genetic Variation
Author
Wishart, Graham
Author
Rowell, David M.
text
Records of the Australian Museum
2008
2008-06-25
60
1
45
86
http://dx.doi.org/10.3853/j.0067-1975.60.2008.1495
journal article
10.3853/j.0067-1975.60.2008.1495
2201-4349
5240070
Misgolas yorkmainae
n.sp.
Figs 7A–J
,
24A
Type material
.
HOLOTYPE
?
,
AM
KS38616
,
Kerewong State Forest
, nr
Lorne
, (
31°36'S
152°34'E
),
15 July 1979
,
pit fall trap
field
No.
1600,
D. Milledge
.
PARATYPES
??
:
AM
KS1553
,
Kerewong
SF, nr
Lorne
, site 108,
Kendall Forest Management Area
, subtropical rainforest (
31°36'S
152°34'E
),
28 May 1978
,
pit fall trap
field
No.
1167,
D. Milledge
.
AM
KS5419
,
Kerewong
SF, nr
Lorne
, site 108(2), (
31°36'S
152°34'E
),
15 July 1979
,
pit fall trap
field
No.
1597,
D. Milledge
.
AM
KS50020
,
Lorne
SF, nr
Lorne
, site 86(3),
31°35'S
152°37'E
,
15 July 1979
,
pit fall trap
field
No.
1593,
D. Milledge
.
AM
KS86230
, all details as for
paratype
AM
KS5419
.
All
type specimens brittle & easily damaged, possibly first preserved in formalin
.
Selection of other material examined
.
??
,
AM
KS38615
,
Bulga State Forest
,
Bobbin Fire Trail
,
100 m
from
Padmans Road
(
31°37'S
152°10'E
),
620 m
,
4 Feb.–9 Apr. 1993
,
pit fall trap
site 57BM,
M. Gray
,
G. Cassis
.
AM
KS48653
, data as for AM KS38615
.
AM
KS38657
,
Mt. Boss State Forest
, 17 1040,
North Plateau Rd.
at
Plateau Beech Picnic Area
. Small gully near picnic area end of
Plateau Rd.
(
31°10'S
152°19'E
),
4 Feb.–9 Apr. 1993
,
pit fall trap
site 32BG,
M. Gray
,
G. Cassis.
AM
KS38661
,
Mt. Boss State Forest
, 17 1120,
North Plateau Rd.
about
1.5 km
by track from
Plateau Beech Picnic Area
(
31°10'S
152°19'E
),
4 Feb.–9 Apr. 1993
,
pit fall trap
site 32BM,
M. Gray
,
G. Cassis
.
Diagnosis
. In male: large dark tan spiders, carapace length c. 8.3–9.1; retrodorsal surface of metatarsi IV with at least 2 spines (
Fig. 7I
); venter entirely pallid (
Fig. 7J
). Bilateral rows of small spines adjacent to scopula on tarsi I and II; bilateral small clusters of spines near distal edges metatarsi I and II (
Fig. 7E–H
). Palpal bulb (
Fig. 7B,C
) embolic flange with c. 10 minute folds; embolus with prolateral bend near tip, apophysis absent. Retrolateral tibial apophysis abruptly swollen midway, distal tibial apophysis massive, cymbium spines strong, attenuate, sub erect (
Fig. 7A,D
). Female not known.
Description
Male
holotype
(
Fig. 7A–J
).
Size
. Carapace length 8.82, width 7.35. Abdomen length 7.64, width 5.00.
Colour
. In alcohol limbs and carapace dark tan; dark brown smudges absent from lateral limb surfaces. Abdomen dorsum dark brown with 6 pallid transverse chevrons. Sparse fine pallid hairs most concentrated on carapace near perimeter & extending onto coxae.Venter entirely pallid.
Carapace
. Edge fringed with many long black bristles which extend onto a raised post foveal surface. Line of c. 9 (some rubbed off) median bristles on caput arch. Group of 6 bristles between PME, 7 bristles on clypeus. Area below clypeus on pleuron membrane not chitinous, without setae. Fovea width 1.43, deep, slightly procurved, edges recurved; posterior wall centrally divided by two pitted intrusions.
Eyes
. Raised on mound. Area immediately adjacent to eyes black; anterior width 1.56, posterior width 1.48, length 0.96, width/length ratio 1.63. Line joining posterior edges of
ALE
transects anterior third of AME. Posterior row recurved in front and behind.
Chelicerae
. Rastellum row of 6(6) strong spines, few smaller spines retreat along pd edges. Intercheliceral tumescence not examined. Fang groove with 9(8) promarginal teeth & 12(12) smaller retromarginal/intermediate row teeth.
Labium
. Bulbous, raised, length 0.87, width 1.05. Labio-sternal suture broad, medially divided.
Maxillae
c. 4–5 (4–5) anteroental basally thickened setae in lieu of developed cuspules.
Sternum
. Length 4.97, width 3.75. Sigilla, anterior pairs small, round, posterior pair larger, ovate, all their own diameter from margin.
Legs
. Tibia I with distal bifid apophysis; both processes “stump-like” with 2(2) short blunt spines.
palp I II III IV femur 4.78 8.53 8.04 6.88 8.63 patella 2.21 4.41 4.11 3.32 4.11 tibia 4.35 6.276.14 4.35 7.84 metatarsus — 6.96 6.32 5.89 8.13 tarsus 2.03 3.52 3.44 3.44 4.05 total 13.37 29.69 28.05 23.88 32.76
Palp
(
Fig. 7A,D
). Cymbium with 37(33) strong, attenuate, sub-erect spines covering anterior half of d surface. RTA abruptly swollen midway, d and rd surfaces covered with short pointed spines; rv edge of tibial excavation and straight massive DTA suspends brush of longer spines. TEM prominent, pallid, adjacent to RTA; TET not seen.
Bulb
. (
Fig. 7B,C
). Embolic rl flange with c. 10 minute folds, margin distally sharply convex curved under embolus as if twisted; embolus with pl bend near tip, apophysis absent.
Scopula
. Complete on tarsi I and II; small remnants on metatarsi I and II; absent on legs III and IV.
Trichobothria
. Palp: tarsi 11, tibia pd 6, rd 6. Leg I: tarsi 13, metatarsi 12, tibia pd 6, rd 6. Leg II: tarsi 12, metatarsi 11, tibia pd 7, rd 7. Leg III: tarsi 12, metatarsi 12, tibia pd 7, rd 7. Leg IV: tarsi 12, metatarsi 13, tibia pd 9, rd 8.
Leg spination
. Leg I (
Fig. 7E,F
): tarsi bilateral rows small spines adjacent to scopula pv 12, rv 16; metatarsi apical pv cluster of 12, rv band of 18; tibia pv 011223. Leg II (
Fig. 7G,H
): tarsi bilateral rows small spines adjacent to scopula pv 16, rv 23; metatarsi pv 011cluster of 9, rv band of 11 including apical cluster of 8, pd 010; tibia v 0111112. Leg III: tarsi v 62; metatarsi v 21, pd 011110, rd 010110; tibia pl 0110, rl 0110; patella pd 2. Leg IV (
Fig. 7I
): tarsi v 48; metatarsi v 25, rd 001010; tibia v 8, rl 01110.
Abdomen
. Dorsum covered with long brown bristles and underlying fine pallid hairs. Venter covered by short brown bristles.
Fig. 7.
Misgolas yorkmainae
n.sp.
A–J
?
, holotype AM KS38616. (
A
), right palp retrolateral. (
B,C
), right bulb:
B
, dorsal;
C
, prolateral. (
D
), left cymbium dorsal. (
E,F
), right tarsus and metatarsus I:
E
, prolateral;
F
, retrolateral. (
G,H
), right tarsus and metatarsus II:
G
, prolateral;
H
, retrolateral. (
I
), right tarsus and metatarsus IV retrolateral. (
J
), venter.
Etymology
. The species is named in recognition of Dr Barbara
York
Main, doyenne of Australian trapdoor spiders.
Distribution and natural history
. (
Fig. 24A
) This species is known only from the
type
locality, the Kerewong State Forest, near the town of Lorne, N.S.W. The burrow is unknown.