Some new Hydroidomedusa (Cnidaria) from the northern South China Sea
Author
Wang, Xuefeng
College of Fisheries, Guangdong Ocean University, Zhanjiang 524088, China
Author
Lin, Kun
College of Fisheries, Guangdong Ocean University, Zhanjiang 524088, China
Author
Xu, Zhenzu
College of Ocean and Earth Sciences, Xiamen University, Xiamen 361102, China Corresponding author, E-mail: guodh @ xmu. edu. cn
Author
Guo, Donghui
College of Ocean and Earth Sciences, Xiamen University, Xiamen 361102, China Corresponding author, E-mail: guodh @ xmu. edu. cn
Author
Huang, Jiaqi
College of Ocean and Earth Sciences, Xiamen University, Xiamen 361102, China Corresponding author, E-mail: guodh @ xmu. edu. cn
text
Zoological Systematics
2019
44
3
191
205
journal article
10.11865/zs.201923
2095-6827
5366458
DC69BC63-5B4D-4851-9A40-715353079BFC
Family
Corymorphidae Allman, 1872
Corymorphidae
Bouillon
et al.
, 2006: 226–227
;
Schuchert, 2010: 377–378
;
Xu
et al.
, 2014: 366–367
.
Diagnosis. The family is diagnosed by: medusa dome-shaped or with apical projection, no exumbrellar cnidocyst tracks, but sometimes long abaxial spurs of marginal bulbs; manubrium mostly not extending beyond umbrella margin, tubular to fusiform, mouth simple, circular; either only one fully developed marginal tentacles, or one fully developed tentacle and three shorter ones differing also in structure; four radial canals and four marginal bulbs present, ocelli lacking; gonad undivided surrounding all length of manubrium and exceptionally also in sac-like processes of manubrium (
Gotoea
).
Remarks. The taxonomic history and synonymies see
Calder (1988)
,
Petersen (1990)
, Pagès & Bouillon (1997), Brinckmann-Voss &
Arai (1998)
,
Bouillon
et al.
(2006)
,
Schuchert (2010)
and
Xu
et al.
(2014)
.
The family comprises 10 medusa genera:
Corymorpha
M Sars, 1835
,
Gosta
Huang, Xu & Lin, 2012,
Eugotoea
Margulis, 1989,
Euphysora
Maas, 1905
,
Gotoea
Uchida, 1927
,
Mayeri
Xu, Huang & Guo, 2012
,
Octovannuccia
Xu, Huang & Lin, 2010
,
Paraeuphysilla
Xu, Huang & Guo, 2011
,
Paragotoea
Kramp, 1942
and
Yakovia
Margulis, 1989 (
Bouillon
et al.
, 2006
;
Xu
et al.
, 2010
,
2014
;
Wang
et al.
, 2011
;
Huang
et al.
, 2012
), of which 8 genera are reported in
China
seas.
Key to the known genera (medusa) of family
Corymorphidae
.
1. With 8 radial canals..............................................................................................................................................................................
2
With 4 radial canals.............................................................................................................................................................................. 3
2. With one developed marginal tentacle, ending in a large knob of cnidocyst, and three small marginal bulbs ....................................... ............................................................................................................................................
Octovannuccia
Xu, Huang & Lin, 2010
*
With 4 equally developed tentacles with numerous adaxial transverse cnidocyst claps and 1 small terminal cluster............................ ..........................................................................................................................................
Paraeuphysilla
Xu, Huang & Guo, 2011
*
3. With 1 fully developed marginal tentacle............................................................................................................................................. 4 Usually with 3 short or rudimentary marginal tentacles and one long, full developed marginal tentacle, different in structure ............ .......................................................................................................................................................................
Euphysora
Maas, 1905
*
4. Exumbrella with longitudinal grooves or ridges and ribs.....................................................................................................................5 Exumbrella with uniform surface......................................................................................................................................................... 6
5. Exumbrella divided in 4 proximal leaf-shaped facets separated by 4 longitudinal large and deep grooves; umbrella without marginal bulb; one developed hollow marginal tentacle; without mesenteries ...........................................................
Eugotoea
Margulis, 1989
Exumbrella with 4 longitudinal ridges and ribs; umbrella with 3 marginal bulbs; with one well developed solid tentacle; with mesenteries..........................................................................................................................................
Costa
Huang,
Xu
&
Lin
, 2012
*
6. Umbrella margin slightly oblique to vertical axis; one developed marginal tentacle, short and thick, ending in a large knob of cnidocysts and three small marginal bulbs .......................................................................................
Mayer
Xu, Huang & Guo, 2012
*
Umbrella margin at right angle to vertical axis ....................................................................................................................................7
7. Gonads on manubrium and on 4 sausages like interradial manubrium pouches ................................................
Gotoea
Uchida, 1927
*
Gonads simple; manubrium without interradial pouches .....................................................................................................................8
8. Fully developed marginal tentacle slender, long, moniliform; pointed apical process .............................
Corymorpha
M. Sars, 1835
*
Fully developed marginal tentacle ending in single cnidocyst capitation or in cluster of cnidocyst capitation; no pointed apical process ............................................................................................................................................................................................................9
9. Fully developed marginal tentacle with a terminal cnidocyst knob; with one tentacle and three non-tentacular bulbs.......................... ...................................................................................................................................................................
Paragotoea
Kramp, 1942
*
Fully developed marginal tentacle with terminal ramifications ending in numerous cnidocyst cluster, with only one marginal bulb bearing tentacle ...............................................................................................................................................
Yakovia
Margulis, 1989
*
Present in
China
Seas