Three new species of Trigonospila Pokorny (Diptera: Tachinidae), from Area de Conservacion Guanacaste, northwestern Costa Rica, with a key for their identification Author Fleming, AJ Author Wood, D. Monty Author Janzen, Daniel H Author Hallwachs, Winnie Author Smith, M. Alex text Biodiversity Data Journal 2015 3 4595 4595 http://dx.doi.org/10.3897/BDJ.3.e4595 journal article http://dx.doi.org/10.3897/BDJ.3.e4595 1314-2828--4595 Trigonospila uniformis Fleming & Wood sp. n. Materials Type status: Holotype . Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0035709 ; recordedBy: D.H. Janzen & W. Hallwachs ; individualID: DHJPAR0035709; individualCount: 1 ; sex: M ; lifeStage: adult ; preparations: pinned; otherCatalogNumbers: 09-SRNP-44688; Taxon: scientificName: Trigonospilauniformis; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Trigonospila; specificEpithet: uniformis; scientificNameAuthorship: Fleming & Wood, 2015; Location: continent: Central America; country: Costa Rica ; countryCode: CR; stateProvince: Alajuela; county: Area de Conservacion Guanacaste; locality: Sector Rincon Rain Forest ; verbatimLocality: Estacion Llanura; verbatimElevation: 135 ; verbatimLatitude: 10.933; verbatimLongitude: -85.253; verbatimCoordinateSystem: Decimal; decimalLatitude: 10.933 ; decimalLongitude: -85.253 ; Identification: identifiedBy: AJ Fleming ; dateIdentified: 2015; Event: samplingProtocol: reared from caterpillar of Stenoma Janzen44 (Elachistidae) ; verbatimEventDate: Jul-15-2009 ; Record Level: language: en; institutionCode: CNC ; collectionCode: Insects ; basisOfRecord: Pinned Specimen Type status: Paratype . Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0040672 ; recordedBy: D.H. Janzen & W. Hallwachs ; individualID: DHJPAR0040672; individualCount: 1 ; sex: F ; lifeStage: adult ; preparations: pinned; otherCatalogNumbers: 10-SRNP-75629; Taxon: scientificName: Trigonospilauniformis; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Trigonospila; specificEpithet: uniformis; scientificNameAuthorship: Fleming & Wood, 2015; Location: continent: Central America; country: Costa Rica ; stateProvince: Alajuela; county: Area de Conservacion Guanacaste; locality: Sector Rincon Rain Forest ; verbatimLocality: Quebrada Bambu; verbatimElevation: 109 ; verbatimLatitude: 10.93; verbatimLongitude: -85.252; verbatimCoordinateSystem: Decimal; decimalLatitude: 10.93 ; decimalLongitude: -85.252 ; Identification: identifiedBy: AJ Fleming ; dateIdentified: 2015; Event: samplingProtocol: reared from caterpillar of Antaeotricha spurca (Elachistidae) ; verbatimEventDate: 26-Apr-2010 ; Record Level: language: en; institutionCode: CNC ; collectionCode: Insects ; basisOfRecord: Pinned Specimen Description Male (Fig. 4a, b, c), 9 mm. Head (Fig. 4b): frontal vitta dark black, slightly tapered apically to twice the width of the ocellar triangle, parafrontal 1/2 as wide as frontal vitta; frontal bristles arise no lower than level of first antennal segment; antennae black; frontoorbital plate silvery-gold turning to black apically; parafacial silvery to slightly gold tinged; palpi black gray; gena 1/8 height of head. Thorax (Fig. 4a): yellow when viewed dorsally with four longitudinal black vittae, these appear fused throughout their length with only slight separation apparent; appearing as two indistinct blotches covering 2/3rds of thorax postsuturally; three postsutural dorsocentral bristles; scutellum bearing white or yellowish pruinosity only at apex (occupying less than 1/5th of total area); 3 pairs of scutellar marginal bristles; subapical scutellars widely divergent, lateral scutellars reduced, almost half the length of the subapicals, these closer to apex, than to basal scutellars; legs black. Wings: pale smoky grayish in color, with one bristle arising at the joint between R1 and R2+3. Abdomen (Figs 1a, 4a): abdominal tergites dark velvety black, with bright, yellow bands covering less than 1/3rd of tergal surface arising at the margins of between the abdominal tergites, these bands not wrapping around to the underside; bright yellow bands straddling the margin between tergites ST1+2, T3, and the anterior margin of T4; tergal bands not possessing a sharp mid-dorsal peak rather appearing flat. Female (Fig. 4d, e, f), 4 mm. Head (Fig. 4e): frontal vitta dark tawny, parallel sided apically equal to twice the width of the ocellar triangle, parafrontal 1.5 times as wide as frontal vitta; frontal bristles arise no lower than level of first antennal segment; proclinate orbital bristles present; antennae light black with orange present at base of first flagellomere; frontoorbital plate entirely gold; parafacial narrow, silvery to slightly gold tinged; palpi light gray at base, with orange tips, slightly haired along upper surface; gena 1/10 height of head. Thorax (Fig. 3d): yellow when viewed dorsally with four longitudinal black vittae, these becoming remaining separate postsuturally, appearing as four distinct lines covering just over 1/2 of thorax postsuturally; three postsutural dorsocentral bristles; scutellum bearing white or yellowish pruinosity over half of its area; scutellar bristles similar to males. Wings: pale smoky grayish in color, with one bristle arising at the joint between R1 and R2+3. Abdomen (Figs 2a, 4d): pointed downward apically so as to appear strongly curved; abdominal tergites dark velvety black, with dull, grayish bands covering at least 1/2 of tergal surface; bands flat and with no distinctive mid-dorsal peaks (Fig. 2a); abdominal bands wrapping around to the underside; bright yellow bands straddling the margin between tergites ST1+2, and T3 with yellow-gray color extending up to and beyond insertion point of median marginal bristles on ST1+2; T3 and T4 possessing 1 pair of medial discal bristles, insertion point of abdominal bristles punctuated by a black outline appearing as black spots. Diagnosis Small black and yellow fly, with 4 prominent black stripes on the thorax, these smudging together so that it appears as 2 large thoracic vittae. Males have a black scutellum, straight conical, and apically pointed abdomen, with 3 narrow gold bands interspersed with black wrapping the abdomen, terminating in a black tip. Female abdomen with a strong down-pointing curve abdominal, 3 grayish abdominal bands lacking mid-dorsal point. Etymology From the Latin "uniformis" , for not changing in form or character, in reference to the uniform nature of the pruinose bands on the abdomen. Distribution Costa Rica, ACG, Prov. Alajuela, rain forest, 109-135 m elevation. Ecology Reared from, Elachistidae , Stenoma Janzen44 and Antaeotricha spurca (2 records). One fly larva per caterpillar.