Phylogeny, taxonomy and flower-breeding ecology of the Colocasiomyia cristata species group (Diptera: Drosophilidae), with descriptions of ten new species Author Takano, Kohei Takenaka Author Gao, Jian-Jun Author Hu, Yao-Guang Author Li, Nan-Nan Author Yafuso, Masako Author Suwito, Awit Author Repin, Rimi Author Pungga, Runi Anak Sylvester Author Meleng, Paulus Ak Author Kaliang, Clement Het Author Chong, Lucy Author Toda, Masanori J. text Zootaxa 2021 2021-12-08 5079 1 1 70 journal article 2931 10.11646/zootaxa.5079.1.1 010695ef-a142-42ad-81d7-2e7ed1f7e89e 1175-5326 5766402 2B118B81-7353-4A2C-A892-5466DFC83230 Colocasiomyia cristata species group Colocasiomyia cristata species group: Okada, 1990: 69 ; Sultana et al ., 2006: 694 ; Fartyal et al ., 2013: 768 . Diagnosis (modified from Fartyal et al . 2013 ). Fore tarsomere II with 4 or 5 equal-sized pegs on lower elongation (ch.14-1,2; Fig. 7A,B ). Hypandrium posterolaterally connected to basal portion of phallal sheath through a pair of gonocoxites ( Fig. 11 ). Shared characters. Arista entirely densely pubescent, without branches ( Figs 3C,G , 19A ). Anterior reclinate orbital seta situated outside proclinate orbital seta ( Fig. 3B,F ). Palpus more or less convex in ventrodistal portion, with prominent seta apically and several shorter ones medially to distally. All cibarial sensilla tiny; anterior sensilla irregularly arranged in square or triangle ( Fig. 4 ). Proboscis without a pair of lateral spherical bumps covered with numerous, short, stout setae. Spiracles small. Abdominal sternite II wider than long, somewhat trapezoid or triangular, posteriorly narrowing. Epandrium anteroventrally much elongated, not fused to cercus, without peg-like, stout seta at posteroventral apex ( Fig. 10A,C,F ). Pregonites absent ( Fig. 11 ). Hypandrium more or less quadrate plate-like, as wide as or slightly wider than phallapodeme, lacking setae, posteriorly articulated to tips of epandrial ventral elongations ( Fig. 11 ). Phallapodeme horizontally flat, fused to phallal sheath ( Fig. 11 ). Oviscapt lateral lobes not fused to each other ( Fig. 16J ); ovisensilla all trichoid ( Fig. 12A,C ). Included taxa. Colocasiomyia colocasiae species subgroup; C . cristata subgroup; and C . seminigra (Duda) *. Geographical distributions of all the component species are shown in Fig. 14 . * Due to the insufficient information in the original description by Duda (1923) , it cannot be determined to which subgroup this species from Papua New Guinea is to be assigned.