Basal Cyclorrhapha In Amber From The Cretaceous And Tertiary (Insecta: Diptera), And Their Relationships: Brachycera In Cretaceous Amber Part Ix David A. Grimaldi Author Grimaldi, David A. Division of Invertebrate Zoology American Museum of Natural History, New York text Bulletin of the American Museum of Natural History 2018 2018-10-24 2018 423 1 97 http://www.bioone.org/doi/10.1206/0003-0090-423.1.1 journal article 7631 10.1206/0003-0090-423.1.1 2e886aea-b59a-45a6-aeaa-2427d584a894 0003-0090 4613008 Canadopeza , new genus DIAGNOSIS: (female only). Acrostichals in 2 even rows; 1 large pair dorsocentrals; M forked, slightly asymmetrical, M 2 very faint, almost spectral; cell dm short, dm-cu less than half the distance between M-Cu fork and wing margin; CuA 2 curved; microtrichia over entire surface of wing; legs without dark, bifid scales; metatarsus not inflated. With the exception of the spectral M 2 vein, these features are plesiomorphic for the Platypezidae . TYPE SPECIES : C. biacrosticha new species . ETYMOLOGY: Derived from Canada , the country of origin, and - peza (foot), a common suffix in the generic names of Platypezidae . COMMENTS: The new genus occupies a basal position in Platypezidae (fig. 27), based on the biserial (vs. uniserial) acrostichals, lack of bifid scales on the legs, metarsus not flattened and expanded, the short dm cell, and vein CuA 2 curved. It is interesting to note that the two Cretaceous genera known from Laurasia ( Canada , New Jersey) are very basal stem groups, along with Burmapeza , n. gen. Otherwise, other Creta- ceous genera, in amber from Lebanon and Myanmar , have a more derived position (fig. 27). It is possible that more basal taxa were sustained longer into the Cretaceous at higher paleolatitudes, or were relegated to higher latitudes, similar to what is seen today with Microsania , Melanderomyia , and Platypezina .