A tentative species list of the European herpetofauna (Amphibia and Reptilia) — an update Author Speybroeck, Jeroen Author Beukema, Wouter Author Crochet, Pierre-André text Zootaxa 2010 2492 1 27 journal article 10.5281/zenodo.195659 645ed7da-f689-4d1c-b7e5-ba1d7120b6ee 1175-5326 195659 Testudines or Chelonii Different opinions exist on whether Testudines (e.g. Fritz and Havas 2007 ; Rhodin et al. 2008 ) or Chelonii (e.g. Bour and Dubois 1985 ) should be used to refer to the the order of turtles, tortoises and terrapins. Spinks and Shaffer ( 2009 ) performed a phylogenetic study of the genus Emys based on multiple genes, both mtDNA and nuclear. There is generally no reciprocal monophyly between Emys orbicularis (European Pond Terrapin ) and E. trinacris (Sicilian Pond Terrapin) in their trees based on nuclear genes, providing additional substantiation for rejection of the latter taxon at species level ( Speybroeck and Crochet 2007 ). Fritz et al. ( 2009 ) investigated the mitochondrial phylogeography of the Spur-thighed Tortoise ( Testudo graeca ) from the western parts of the Mediterranean, and recognised T. g. nabeulensis as a valid subspecies for the Tunisian populations, with Sardinian and Sicilian animals belonging to this taxon. In this arrangement, Majorcan and Spanish populations remain treated as T. g. graeca .