World revision of the microbombyliid genus Mythenteles Hall & Evenhuis (Diptera: Mythicomyiidae) Author Evenhuis, Neal L. text Zootaxa 2003 346 1 28 journal article 51254 10.5281/zenodo.157139 0cf8bc38-65c0-45cf-a4d0-2eb229661dff 1175­5326 157139 Mythenteles silus Evenhuis , sp. n. ( Figs. 11 , 22 , 28, 34) DIAGNOSIS. Closest in appearance to M. deemingi , sp. n. by virtue of the short proboscis, but can easily be separated from it by the short second antennal flagellomere (less than 1/3 the length of the first flagellomere; about 3/4 the length of the first flagellomere in M. deemingi ), the anepimeron with a spot of yellow color, and the thick labellum (thin in M. deemingi ). DESCRIPTION. Male . Length: 0.95–1.25 mm . Head . Black; eyes dichoptic, separated at vertex by 1.5 x distance between lateral ocelli; frons depressed medially, yellow to yellow­white; face yellow, tip of oral margin brown; antennae brown; scape minute; pedicel cylindrical, slightly wider than long; first flagellomere ( Fig.11 ) ovoid, length about 1.5 x greatest width; second flagellomere about 1/3 length of first flagellomere, with apical sensillum; occiput and mentum shiny brown; proboscis brown, thick, length slightly longer than head height; labrum sclerotized, stiff, pointed apically, length one­half head height; palpus not evident. Thorax . Mesonotum and scutellum subshining dark brown to black, with scattered dark hairs; remainder of thorax as in M. deemingi with addition of yellow posterodorsal corner of katepimeron. Wing ( Fig. 22 ). Hyaline; veins brown; costa ends slightly less than halfway between end of R4+5 and M1; vein Sc incomplete, ending at level about halfway between origin of Rs and end of vein R2+3; origin of vein R2+3 almost perpendicular to Rs; Rs evanescent at connection with R1; R4+5 straight to wing margin, not curved, ending in costa at level before end of vein M1; vein M1 curved basally, then straight toward wing margin; M2 straight to wing margin; crossvein dm­cu closing cell dm present; CuA2 thick along cell dm, thin on apical 2/3 to wing margin; A1 straight to wing margin, not curved or sinuous; fringe of hair on posterior margin of wing rather long. Abdomen . Dorsum dark brown, with scattered dark hairs; yellow laterally on tergites II–VII, sometimes extending medially as posterior fasciae, especially so on apical tergites; venter paler brown than dorsum, with yellow on posterior portion of segments. Genitalia (Fig. 28). Gonocoxae higher than wide, ovoid, with finger­like projections on apicoventral surface, with X­shaped sclerotization in dorsal view; gonostylus relatively large, ca. 1/3 length of gonocoxa, bifid apically; epiphallus long, thin, with long thin apodemes connected to aedeagal sheath, upper portion vase­like in ventral view, with bifid apex; aedeagus long, thin, with small aedeagal bulb, aedeagal apodeme shoe­shaped in lateral view, with small lateral vanes; epandrium subrectangular in lateral view, wider than high, with curved pseudosurstylus, cerci small, slightly exerted. Female . Same as male except as follows: female genitalia ( Fig. 34 ) with vaginal furca U­shaped, with apical processes interrupted medially, medial processes perpendicular to lateral processes; spermathecal reservoir ovate­conical, sclerotized brown; apical spermathecal duct long, thin; sperm pump slightly longer than vaginal furca, flared apically; basal valve not evident, basal duct short, membranous, leading to common duct. Types . Holotype male and 8 male and female paratypes from UNITED STATES : California : San Bernardino Co: 4 mi . [ 6.4 km ] E of Cushenbury Springs, 7.v.1980 , J.C. Hall. Paratypes . MEXICO : Baja California : 1 male , 1 female , Sierra San Pedro Mártir, trail La Joya to La Zanja, 10.vi.1953 , P.H. Arnaud, Jr. (CAS). UNITED STATES : Arizona : [Coconino Co:] 1 male , Williams, 7.vi. , Barber & Schwarz (USNM). [Cochise Co:] 1 female , Huachuca Mts, Miller Cyn, 4.v.1948 , A.L. Melander (USNM). Pima Co: 3 females , Organ Pipe Cactus National Monument, Williams Spr, 15–17.ii.1970 , Arnaud (CAS). California : 3 males , 9 females , topotypic, collected with types (BPBM, CAS, UCR, USNM). Alpine Co: 1 male , 1.5 mi . [ 2.42 km ] NE of Red Lake, 25.vii.1955 , E.I. Schlinger (UCR). El Dorado Co: 11 females , 2–4 mi [ 3.2–6.4 km ] S of Meyers, 24.vii.1955 , E.I. Schlinger UCR). Riverside Co: 1 male , Aguas Caliente Indian Reservation, Palm Cyn, 24.ii.1970 , Arnaud (CAS); 1 female , 5 mi [ 8.1 km ] E of Elsinore, R.R. Cyn., 13.iv.1965 , C.A. Toschi (UCB); 1 male , 2 females , Elsinore Mts, nr summit on Hwy 74, 24.vi.1964 , E.F. Riek & G.I. Stage (UCB); 1 female , Menifee Val, (hills on w end), 1800 ft [ 548.6 m ], 15–30.vi.1980 , J.D. Pinto (UCR); 2 females , Ortega Hwy, El Cariso Camp, 26.v.1944 , 15.v.1946 , Melander (USNM); 1 female , Palm Springs, 15.iii.1955 , Melander (USNM). San Bernardino Co: 1 female , nr Adelanto, 23.v.1945 , Melander (USNM); 1 female , Barton Flats, 22.vi.1948 , Melander (USNM); 1 male , Cajon, 22.v.1945 , Melander (USNM); 1 female , Cajon Cyn, 9.v.1944 , Melander (UCR); 2 males , 8 females , Camp Angelus, 26–28.v.1947 , Melander (USNM); 1 female , Mill Cr. Camp, 20.v.1945 , Melander (USNM); 1 female , Morongo Val, 20.iv.1944 , Melander (USNM); 1 male , 1 female , Mountain Home, 21.vi.1945 , Melander (USNM); 1 female , Oak Grove, 9.v.1945 , Melander, on white Ceanothus (USNM); 5 males , 15 females , Sheep Cr. Cyn., 24.v.1945 , Melander (USNM); 1 female , South Fork Sta. Ana Riv., 19.vi.1945 , Melander, on white Ceanothus (USNM); 1 female , Upper Sta. Ana Riv., 21.v.1947 , Melander (USNM); 4 females , Wrightwood, 22.v.1954 , Melander (USNM). San Diego Co.: 1 male , Borrego, 5.iv.1949 , W.W. Wirth (USNM). Nevada : 1 female , Charleston Mts, Kyle Cyn, 5600 ft [ 1706.9 m ], 4.vi.1944 , P.H. Timberlake (UCR); 2 females , Mercury, Nevada Test Site, 6.v.1961 , BYU­Atomic Energy Commission Collection (USNM). Holotype male and paratype female deposited in CAS. Other paratypes in USNM, UCR, and BPBM. Remarks . As in propleuralis , there are numerous individuals of this species that vary in the presence, partial presence, or absence of crossvein dm­cu closing cell dm. Most of the specimens included here in the type series of silus were previously a part of the paratype series of Mythicomyia mutabilis Melander. It should be noted that one specimen with a mutabilis paratype label examined during this study is actually a species (undetermined) of Mythicomyia ( Mythicomyia ) . Etymology . The specific epithet derives from the Latin silus = pug­nosed; referring to the short, thick labellum and normally pale­colored proboscis. Distribution . Mountainous California to Nevada and Arizona and into adjacent areas in Baja California in Mexico . Further collecting may also show it to occur in Sonora, Mexico .